Bulletins of American paleontology (Bull. Am. paleontol.) Vol 248087193839

r
-^.y.

\5

BIMJ.ETINS
OI'

\

M R CA N
V.

1

^-\LlOXTOLOG^

VOL. XXIV

3i^

0> vJ^C-^-^O)

E

m

llMACA. N. Y.
IT.

S.

A.



CONTENTS OF VOLUME XXIV
Bulletin No.

Plates

80.

Nomenclatorial notes on Eocene Mollusca
By Katherine VanWinkle Palmer

81.

Devonian crinoids from the Mackenzie River Basin,
N. W. T., Canada
By Winifred Goldring
1-2
The correlation of certain Devonian faunas of eastern
and western Gaspe
By E. M. Kindle
3-4
Devonian Bryozoa of Gaspe
5-6
By Madeliene A. Fritz
A Devonian fauna from Colombia
By Kenneth E. Caster; Including Stratigraphic

Notes
By Axel A. Olsson
7-20
Notes on Cypraea heilprini Dall and Cypraea chilona
Dall with new species from the Pliocene of Costa

82.

82A.
83.

84.

Rica
By William Marcus Ingram
85.

86.

Pages
1-7

8-34

35-86

87-100

101-318

21

319-326

22

327-340

23-26

341-359

of Pennsylvanian crinoids from the vicinity
of Bartlesville, Oklahoma
27-29
By Harrell L. Strimple

359-386

New

_

from the Miocene of the Dominican Republic and Panama, with a survey of the
Miocene species of the Dominican Republic
By William Marcus Ingram
Reprint of Conrad's Jackson Eocene fossils as described and illustrated in the Philadelphia Academy
fossil Cypraeidae

of Natural Sciences, Proceedings for 1855, pp. 257-63

and Wailes' Report on the Agriculture and Geology
of Mississippi, 1854, pis.
87.

XIV-XVII

.-..

A group

.

Index

_

.

.

_



BULLETINS
AMERICAN

PALEONTOLOGY
VOL. XXIV

NUMBER
SO

Ithaca,

U.

New
S.

York,

A.

f^.



BUI
OF

AMERICAN PAI^BONTOLOGY
Vol. 24

No. 80

Nomenclatorial Notes On Eocene Mollusca

By
Katherine

July

Ithaca,

VanW. Palmer

I,

1938

New

York,

U. S. A.



NOAIENCLATORIAL NOTES ON EOCENE MOLLUSCA
By

Katherine \'an Winkle I'almer

At the time

of the puljHcatiori of the munoii-rapli

on

tlie

Clai-

bornian fauna^ the reference showing Ccclatitra Conrad, i8()5was preoccupic(h had not been found. Since that time the author
has located the citation wliich Conrad i;rol)ably liad in nhnd when
he changed the

name

of the

genus to Actcioncuuir

Tlie confusion in connection

discussed in detail

Claiborne and

will

))}

with these two names has been

the writer in the

work on the fa.una of the
Conrad himself used

not be repeated here.
iS53'* for a

Ccclatiira previousl_\- in

Naiad.

Actccoiicina

fore a substitute r.ame for Ccrlatura C^inrad,

lem

of

its

genotype

falls in that

which was suggested on

p.

categorw

i
The one

is

there-

and the probinterpretation

156 of the Claiborne work

ma_\-

be

eliminated.

Aldrichia Palmer''
insects aiid

named

X'aughan"

is

])reoccupied

in corals.

1)\'

.ihlricliia

Aldrichia Palmer

CrKpiillett''
is

in

herein re-

Tiinothia.

Attention

is

called to the reference by J.

W.

dates of the publication of the various parts of

"Histoire Naturelle des Mollusques Terrestres

Taxlor" on the

Moquin-Tandon's
et

Muviatiles de

iPalmer, K. V. W., Bull. Aiiicr. l^al., vol. 7, No. :V1, p. 154, 1S>:!7.
2Conrad, T. A., Amer. Jour, ("oiu'li.. vol. I, pp. 28, .'iS, 1865.
aConrad, T. A., ibid. y. 147.
4Coiirad, T. A., Acad. Nat. Sci., I'hila., Proc. vol. 6, p. 268. 185;i.
sPaliner, K. V. W., Bull., Amer. Pal., vol. 7, No. ;;2, p. 262, 1937.
sCoquillett, D. W., Trans. Amer. Entomol. Soc, vol. XXI, p. 9:5, 1894.
-Vaughr.n. T. W.. U. S. Geol. Sur., Men. vol. XXXIX, ]>. 70, 1900; Proc.
Biol. See. Wash., vol. XYI, p. 101. 1903.
Aldrichia Vaughan renamed.
sTaylor, J. W., Proc. Mai. Soc. London, vol. 6, p. 186, 190-1.

ikYti w«


4

Bulletin 80

4

were issued April 12, May 4, August i,
September 10, 1855 January 2, April 9, 1856 respectively. These
Conrad, Janudates definitely establish the priorit>- of PapilUna

The

France."

six parts
;

PapilUna AIoquin-Tandon, 1855.
Aluseum pointed
F. Stearns MacXeil, United States National
a t}pe for Cladesignated
out to the writer that Herrmannsen"

ary, 1855^ over

HerrSwainson, 1840 previous to other designations.
of
discussion
mannsen's statement of type was overlooked in the

z'ilithes

by the author^^ and the work of
The problem of Clavilithes Swainson.

of the Claibornian Clavilitlies

Grabau'- was followed.

1840" begins vritli Clavella Swainson, 1835^* for which it was a
name. How^ever, it does not seem that with the exception of one genus (p. 7,"Trochilea, type Trochus pilens. Auct.")

Swainson's "Elements of Modern Conchology" can be used for
He did not mention any
generic names without difficulty.
substitvite

names or references (exception p. 14, Mitreola, reference
The generic names are descriptions without species and
would therefore require special ruling as under Int. Rules Zool.
Nomenclature Opinion No. 46. In case of Clavella, the description reads, p. 20, "The genus Fusus, again, has no plaits it is so

specific

given).

;

closely allied to the fossil

that there can be

group";

no

p. 21,

plaits,
It

genus Clavella (here

no doubt

of

"Clavella Sw.

its

now

first

defined)

entering within the limits of this

Fuciform [fusiform]

but the tip of the spire enlarged.

;

channel long;

Fossil only."

appears that even though Clavilithes Swainson, 1840 was a

substitute

name

no
must take its type from the species mentioned
own description and the type so established becomes
for Clavella Swainson, 1835, since Clavella has

species, Clavilithes

under

its

the type of both genera (Int. Rules Zool.

Nomen., Art.

30, f.).

sConrad, T. A., Acad. Nat. Sci., Phila. Proc. vol. 7, p. 262, 1855 Palmer,
K. V. W., Bull. Amer. Pal. vol. 7, No. 32, p. 363, 1937.
loHerrmannsen, A. N., Indicis Generum Malaeozoorum, vol. I, p. 246,
;

1S46.

iiPalmer, K. V. W., ibid, p. 356.
i2Grabau, A. W., Smith. Misc. Coll., vol. XLIV, p. 104, 1904.
isSwainson, Wm., A Treatise on Malacology etc., p. 304, 1840.

i^-Swainson,

Wm., Elements

of

Modern

Coneholog^-, pp. 20, 21, 1835.


NoMENCLATUIilAL NOTES

:

K. V. W. PaLMER

Herrmannsen designated Fnsus noce (Chem.). This is a valid
designation as F. nocc was listed in Swainson's original descripThe characters of F. noce disagree with
tion of Clavilithes, 1840.
the original description of Clavella because F. nocc (Chem.) Lam.
has plications on the columella during part of

its life

history, such

Mr. A. Wrigley, England^^ who

disappearing with maturity.

was consulted in this matter has expressed the opinion that such
noncomformity eliminates the use of F. noce (Chem.) Lam. as

a

genotype of Clavilithes.
also used.
of the

'

The

vagueness of

Losing F. noce

This

is

the reasoning w^hich Grabau^*^

writer does not favor this interpretation because

type of Clavella Swainson, 1835.

tlie

Lam.

as the genotype of Clavilithes

Swainson

causes a change in the current idea of the genus as well as a
conflict

with the subsequently

Lam.
Rhopalithes Grabau

named genus Rhopalithes Grabau/'

type F. noce

becomes

synonymous

with

Clavilithes

Swainson, 1840 and the forms of Clavilithes typified by C. parisiensis (Mayer-Eymar)=rC. longcevus (Desh.) non Solander are
without a generic or subgeneric name.

It is in this last

nonplicate

group that the Claibornian species belong.
However, there is
some doubt that Clavilithes needs to be separated on the character
of the columellar plications,

which do have

particularly

plications in the

young

when

stages of

those specimens

growth

lose

them

in the adult.

The presence or absence

of columellar plications

cannot always

be taken as a factor of generic differentiation in the gastropods.

A

example

genus including plicate and nonplicate
Eocene genus Mazzalina Conrad, i860
= (Bulbifiisus Conrad, 1865).^* Conrad made two genera on
characters which are now known to be only specific and in some
typical

shells

cases

is

of a

the Claibornian

may

not even be specific.

isPersonal letter. May 7, 1938.
i^Grabau, A. W., ibid, p. 104.
^
'Grabau, A. W., ibid, p. 135.
isHarris, G. D., Ark. Geol. Sur., Ann. Kept. State Geol.,
1892; Palmer, K. V. W., ibid, p. 349.

vol.

II, p.

165.


6

Bulletin 80

Q

The

prol^lem of Clarillthes

burdened

with

thorough work.
Swainson, 1840,

more

is

names

already too complicated to be

without

further

complete

and

Until then, the author prefers to use ClavUithes
{=Rlwpalithcs Grabau, 1904) genotxpe by sub-

sequent designation,

Herrmannsen, 1846,

Fitsits

(Chem.)

noce

Lam. to include the nonplicate forms of C. parisiensis (MayerEymar) Wrigley's'^ criticism of Grabau's study of the phylogen>'
.

of ClavUithes

shows that additional investigation must be made

on the group before a satisfactory conclusion is reached.
Since needless time is wasted in searching for Eulimella Forbes,
1:21846, as given b_\- authors-" it seems worthwhile to reiterate
dale's^^ affirmation that

no

sLich reference exists.

One

will

fmd

is an article by Forbes in the Ann. Mag. Nat. Hist., vol.
412 but the genus Eulimella is no where mentioned in

that there

XIV,

p.

The date of the reference is 1844 instead of 1846.
Malaconchologists as H. and A. Adams, Fischer, Bucquoy, Dautzenberg and DoUfuss, Tryon, Sacco, Cossmann and many other

the article.

standard authors continued the error in their work. According
was the first to mention EtiUmella in liter-

to Iredale, Jeffreys-'

ature.

Jeffreys

was followed by Gray-^

gave the name and selected a type.

in the

same' year.

Gray

Iredale suggested that the

name Eulimella

as ascribed to Forbes was a manuscript name.
Such a supposition appears reasonable for particularly Jeffreys
was assisting Forbes^* in the work on the British Mollusca. Iredale prefers to give Gray credit for publishing the genus. Thiele-^
in his Handbuch assigns the genus to Gray.
However, it seems
to the

author that Jeffreys' reference of Eulimella

but with no designation of t}pe.

He

is

states "Eulimella

legitimate

(Forbes")

loWrigley, A. G., Proc. Mai. Soc. London, vol. XVII, pp. 222, 234-237,
1927.
soDall, W. H. and Bartseh, P., Bull. IT. S. Nat. Mus., No. 68, pp. 10, 17,
See Iredale, T., The Nautilus, vol. XXIV, No. 5, p. 53, 1910 for
1909.
previous lefereiiees.
2ilredale, I., ihid, p. 53.
aaJcffreys, J. G., Ann. Mag-. Nat. Hist., vol. XIX, p. 311, May, 1847.

23Gray, J. E., Proc. Zool. Soc. London, pt. XV, p. 160, Nov. 9 (read),

1847.
SAjeffreys, J. G., Hid, p. 309.


NOMENCLATORIAL NOTES

c;-c::sula, I\Ial.

Hi-t., vcl.

and Conch.

XiW

p.

412

.

.

J.
.

E.

MacAndrci, Forbes

Ann. Nat.

in

.

"

Eiilinnila (jracilis

K. V. W. PaLMER

:

(Followed

l\v

a description of this

last species.)

Authors-"

l;elicve that the E.

Andrei Forbes and
(Scacchi)

is

EuUuiclla and

genus

the

the species
it

is

crassida Jeffreys equals E.

same as E.

Tujrbes.

scUlcc (Scacchi).

Gra_\'

Vv-hich

the species which

assigned to

is

species

is

is

E.

Macscillar

used as the type of his

commonly used when

the

Fortunately therefore the same

can be designated as the type of EuUmclla

(Forbes)

Jeffreys and the established characters of the genus need not be
disturbed.
is

To

bring this about, in case the authority of the genus

granted Jeffrc}s, the genot\pe of EuUmella (lu^rbes) Jeffreys,

1847

herein designated as E.

^^

Andrei

(

crassula

Forbes) =/:. sciUcc (Scacchi).

and Scandinavia.

(Jeffreys)

Recent.

Fliocene and Pleistocene of

=£. Mac-

Great Britian

Ital}-

and

Sicil}-.

2'''T!!i?le, J., II:iii(ll)i:e!i diT Sy:itcmati;:'c'.:on Wcichtievkundo, pt. !, p. LD'.O,
1929.
2GForbes, E. and Hanley, S., Hist. Bntinh Moll., vol. Ill, j). ;;;i9. 1S')1
Dall, W. H. and Bartfch, P., U. S. Nat. Mus., Bull. No. CS, p. 10, l::; ;).
;

Paleontological Eepearch Institltion

May

25, 1938.



BULLETINS
AMERICAN

PALEONTOLOGY
VOL. XXIV

NUMBER
81

Ithaca,

U.

New
vS.

York,

A.



BUI,I,BTINS

OF

AMERICAN PAI,EONTOI,OGY
Vol. 24

No. 8 c

Devonian Crinoids from the Mackenzie River Basin,
N.

W.

T.,

Canada

By

Winifred Goldring

August

Ithaca,

U.

1938

15

New
S.

York.

A.



DEVONIAN CRINOIDS FROM THE MACKENZIE RIVER
BASIN, N. W. T., CANADA
By

Winifred Goldring

New York

State

Museum, Albany, N. Y.

from the Great Slave Lake
Northwest Territories, Canada,
was submitted to the writer for study by Doctor E. M. Kindle,
Victoria Memorial Museum, Ottawa.
Of this collection he
writes, "The horizon is probably not far from that rej^resented by
Ivecently a collection of crinoids

Mackenzie River

region,

basin.

two of our reports".
two new species Melocrinus borealis
and M. canadensis collected by E. J. Whittaker from the Hay
River section, the former below the Alexandra falls and the latThe Hay River section from
ter above the falls (ref. cit., p. 17).
which the crinoids were obtained was studied by Kindle who re-

crinoids described and figured by Springer in

In 1921 Springer described

4) the beds to the Upper Devonian, having found
a characteristic Portage fauna in the Simpson shale below the

ferred (1919,

p.

Melocrinus borealis

is

represented in the present collection from locality 7005, bed

h,

strata

from which the crinoids came.

Lady Evelyn

Falls section of the

locality 7300, the
fully referred to

Kakisa River; a few plates from

gorge section of the Redknife River are doubt-

M.

canadensis.

Springer relates M. borealis to

form described by Rowley (1893, p. 303;
1894, pp. 151, 153) from shales considered of Middle Devonian
later authorities of
(Hamilton) age by early geologists and
younger age {see Keyes 1894, p. 43; 1902, p. 271-273; Greger

M.

fersns, a Missouri

1)_\-

374; Sehuchert 1903a, p. 143, 1903b, p. 545; Weller,
1909, p. 264; Branson 1923, pp. 44-46). Springer concludes, "it
is clear that the fossils of the Missouri and Mackenzie Devonian
belong to the same palaeontological province, and are of approx^
p.

imatel}' the

same age"

(ref. cit. p. 15).


12

Bulletin 81

In his later paper (1926) Springer adds three new species of
Mclocrinus: M. kindlei and M. mackenzic from the coral reef in
limestone above the horizon of the Simpson shale. Root River
section and .1/. zvhiftakcri from the beds at least 300 feet above
;

No specimens of
and only a single specimen
inackeii::ic was found and this in the

shale in the Trout River section.

the

Simpson

A[.

kindici appear in the collection,

that can be referred to

.1/.

crinoid bed at the upper falls of the Redknife River.

specimens from the Redknife River section,

at the

new

upper

falls,

M.

species,

At

in the

least three

crinoid bed

Three
have been referred to M. zvhiifakeri.
M. sidcosutura and M. humei, are

subfilistriatiis,

here added from the Redknife River crinoid bed.

Springer also describes

(p.

132)

one species of H'exacrimis

H. hujnei) but Melocrimis is the only camerate genus represented to anv extent in the three collections and it is so far represented
Except for M. borealis which shows clo'se relab\' eight species.
tionship to species of Iowa, Missouri and Wisconsin, as pointed
(

;

out

b\-

Springer (1926,

127), the species of Melocrimis are "not.

p.

And

only thoroughly distinct from that, but also from each other.
the interesting thing about

them from

that in the characters by

which they

all other known American species,
M. mackcnzie and M. zvhittakeri']

a geological point of

the three

.

.

species [Af. kindlei,

exhibit a tendency to an
is

not observed

the abundant species of the Eifel limestone of the

which developed

member

nian (or lower)
koiiiiicki,

M.

in certain species
of the

Jiieroglyphicus

is

from

differ so completely

metrical construction of the calyx which

ian, but

view

asym-

among

Middle Devon-

belonging to the Fras-

Upper Devonian

in

Belgium

\_M.

(non Goldf.) Fraipont=Af. dewal-

von Koenen, M. henedeni and M. }iiespilijoniiis'\" (see also
Von Koenen, 1886). M. borealis has a
p.
similar, though less marked tendenc\' to as\'mmetr\' of the calyx,
qitci

129; Fraipont, 1883;

and

new species described here it as well-marked as
The single specimen of Hexacriuus in the collec-

in the three

in the others.

tion studied

alent ^liddle

by Springer (1926) represents "another very prevDevonian genus in the Eifel, but rare in x\merica.

Mackenzie Ckinoids: Goldring

... of a type complete!}- different from that of the
is

but which

TLifel,

Upper Devonian rocks

also represented in the

Belgium"

of

(p. 127).

In the collection untler consideration inadunate species of crin-

Lake Kakisa, one

oids are represented from

the south shore, west end

;

and eight miles above the mouth
Redknife River,

in the

half mile

back from

the gorge section of the Bouvier River
;

above the upper

gorge section and

falls

of the

at the third chute.

A

single specimen representing the Flexibilia was collected from
the crinoid bed near the base of the coral zone, Jean Marie River,

was found loose in the Trout River section, at the
foot of the long heavy rapid, one half mile below the lower cascade.

Only portions of the vertical side of two arms of the starfish are preserved, and not in very good condition, so that this
and a

startish

species has not been placed.

The

collections studied by Springer

taker in the
for the

Hay

were made by E.
by G.

River and Trout River sections

Root River

section.

In his later paper

.Springer has incorporated notes by

Mr.

;

(1926,

Whittaker

J.

Whit-

vS.

Hume

p.

and

128)

Mr.

Hume, relative to the stratigraphy of the crinoid-bearing beds.
The present collection, in so far as labelled, was collected by E.
The species of crinoids described in all collections
J. Whittaker.
so far submitted for stud\- are

CAMERATA

Melocrinus borealis Springer

M. canadensis Springer
M. kindlei Springer
M. mackenzie Springer
M. whittakeri Springer
M. sulcosutura Goldring
JM. subtilistriatus
?il.

I'.umei

Goldring

Goldring

Hexacrinus humei Springer


i'*

Bulletin 81

6

FLEXIBILIA
Synaptcjcrinus

(

rotunclatus Cioldring

?)

INADUNATA
Undetermined

;

sp.

Decadocrinus spinobrachiatus Goldring
Prininocrinus robustus Goldring

Linocrinus kindlei (ioldring

DESCRIPTION OF SPECIES
CAMERATA
Melocrinus canadensis Springer
Melocrinus canadensis Springer, Geol. Surv., Canada, Bui. 33,

p.

17, pi.

1921.

1, fig. 3,

Melocrinus canadensis was based by Springer upon a single

specimen lacking basal plates and
trix.
falls

onl_\-

from the ma-

parti}' free

In this collection, from the Trout River section at the third
(loc.

6978), are a few radial and interradial plates, partly

separated, which might be referred to this species

and then only

with doubt.

The only
of a larger

description given with the figure

and more robust

ness of which "this form

ton of western

t_\-pe \\'ith

b_\-

that this species

is

in the flat-

is

comparalile with one from the Hamil-

New York

figured by Hall, but ne\'er described,

under the name M. breviradiatus"
figured

is

very low plates,

Hall in 1872

(pi.

i,

(ref. cit., p.

figs.

18,

17).

The form

19) has since been de-

and refigured by the writer ( 1923, p. 127-130, pi. 13, figs, i
There is even more resemblance to Melocrinus clarkei (Hall
Ms) Williams from ^enesee and Portage beds of the Upper De-

scri])ed

2).

(

vonian of western
3-5

;

pi.

14).

The

New York

{see

anal tube, not characteristic of

nor found

in

any

ref. cit., p.

figured specimen of

of the

M.

132-136,

pi.

13, figs.

shows a short
the other species under discussion

specimens in

canadejisis

this collection.


MACKEiVziE Ckinoids: Goldrin'G

15

Melocrhilic whittakeri Springer

Mclocrinus

irhitlfiL-rri Spriii;L;x r,

Mclocrinus

The

stant.

15 miles

300

least

in

holot_\])e

was based l)y Sprinj^er upon three wellwhich the characters are thoroughlv concame from the Trout River section, about

its confluence with the AJa,ckenzie, "from Ijcds at
above the Simpson shale and thought to be s(nne-

feet

M.

tlian the
(

species

ref.

witli

cit.,

M.

]).

borealis horizon of the Ha\- River, l'p])er
In discussing the relationship of

132).

k'uidlci

and

Ri\er section Springer states that

them and

42, pp. 1;!1, l;i2,

above

what higher
Devonian"
this

Canada, Bui.

iriurv.,

Tvliittakcri

specimens

(letined

Ueol.

1926.

14-17,

24, fios.

pi.

others

all

known

]\\-

its

uiackcnrjic

.1/.
it

from the Root

"readily distinguished from

is

marked ovoid contour, and ex-

tremely small column facet, whicli indicates a considerabl\- different type of

column from

The tendenc}
species

is

that of the

at all similar to this

is

genus as generalh- found.

None

usually to a broad base.

of the Belgian

except in asymmetry"

(ref.

cit.).

In the collection submitted by Dr. Kindle are at least three

specimens that might be referred to
san;;e

.1/.

crinoid bed were collected three

but from

t^'hittakcri,

new

species

all

tlie

showing

a

small column facet and two of tliem with elongate ovoid cahx.

These new species

howe\er, easily distinguished from M.

are.

whittakeri.

One
large,

of the

The second
(24

specimens referred to

measuring zy mm.

mm.

bases).

to

to the

this

species

arm bases with

a

is

abnormally

broken basal cup.

s])ecimen has a height for the calyx of 28.2

arm bases)

;

the third 25.5

mm.

{22.2

mm.

to

mm.
arm

All the specimens, therefore, are larger than those de-

scribed by S])ringer

;

and the writer

l)elie\-es

mature forms, as indicated by the character

that they are

of the plates.

more

.Second-

ary thickening of crinoid plates develops in older forms sometimes
with quite striking dift'erence

in

scribes the plates of the calyx as

i)late

characters.

"smooth or

Si)ringer de-

slightly rugose, flat,

with a slight tendency to pitting at the angles, but without convex-


16

Bulletin 81

8

or median elevation either in dorsal cup or tegmen" (p. 131).
In these three specimens the pitting at the corners is well shown.
Thickening of the plates is seen at the margins, and there is a

ity

surrounded by

central raised area or flattened tubercle

In the second

depression due to the thickening at the margins.
plate

largest specimen an occasional

shows

a

a slight

more prominent

central tubercle.

Horizon and

locality.

— From the crinoid bed

at the

upper

falls,

Redknife River, locality 7288.
Melocrinus subtilistriatus

n.

Plate

sp.

1,

figs.

In the collection submitted by Dr. Kindle are a fairly large

1-5

num-

ber of specimens which in the shape of the calyx and the small
facet bear a strong resemblance to

column

25

are of

mm. and
from 15.7 mm. to

average calices measuring between 20.2

high with a width at the

Two

whittakeri.

The specimens

appears to be a smaller species.
size,

M.

arm

bases

particularly large specimens have heights of 24.6

mm.

This

medium
22 mm.
18 mm.
mm. and

All the specimens have asymmetrical, elongate ovoid

below to a very narrow
column facet, and also between
the veiy low tegmen with subcentral anal opening

calices contracting

more or

less strongly

base, with the characteristic small

the rays into

without a tube.

Specimens

in this .collection,

if

seen alone, might give the im-

pression of being varieties of the species, or even different species,

because of the presence in some of faint

stellate

ornamentation,

and more tumid plates. However, the
writer has picked out a series showing the relation of one stage
One specimen shows the "smooth or slightly rugose,
to the next.
flat plates", referred to by Springer (1926, p. 131) in his descripin others of raised ridges

tion of

M.

whittakeri, with pitting at the angles.

shows remnants
ticularly well

Closer inspection

of delicate carinse crossing the suture lines, par-

shown on the

A

radials, first

primibrachs and primary

second specimen shows beautifully a delicate
ornamentation of groups of two or three fine carinae extending

interbrachials.


Mackenzie Okinuids: Goldring

17

from center to center of the hasals, raclials, first primibraclis and
primary interbracliials. Usually the center ridge of each group
is

The higher

stronger.

and interradial

plates of the radial

series

have strong ridges, usually only one running from center to center
with an accomi)anying deeper pitting at the angles.

grew

viduals

x-\s

the indi-

older changes in the character of the ornamentation

took place, well-shown in the selected series of specimens.
thicken

carin;e

with

ridges

into

low ridge following each radial

and with

rugose character

this a

is

thickening process continues the

this

I'he
of

a

is

de-

of

the

the center of the plates, ]iarticularly the i)rimar_\- inter-

b'"achials,

tumid,

Sometimes there
marking the junction

series.

veloped a central blunt tubercle,
cariui'e at

As

development

the

witli the

presence of cariuce indicated,

gixen to the plates.

plates
if

become

at all,

tjuite

only at the

sutures and on the higher interbrachials, especially of the anal

Thickening

interradius.

of the plates at the

margin tends

to de-

velop depressed sutures.

Horizon and

locality.

— The

cotypes are from locality 7288, the

Redknife River. There are other
locality
and from localit}' 7291, bed
from
same
the
specimens
marked c (field No. 267), 34 "lile below the upper falls, Redknife
River; probably also from localit}' 7005, bed h, Lady Evelyn F'alls
crinoid bed

section,

at the

upper

Kakisa River.

Remarks.

—The

specific

striated character of the

rpecies

falls,

from both M.

Melocrinus sulcosutura

name

is

given because of the finely

ornamentation which distinguishes

zvhiffakeri

this

and M. sulcosufura.
Plate

n. sp.

1,

fig.

6

From the same crincjid bed in which M. sitbtilisfriatiis is so
abundant was collected a single somewhat crushed specimen of
the same type, but with enough differences to warrant placing it
new

cup and part of the
from
the
gorge section at the
tegmen are preserved. A fragment
lowest chute, Redknife River (loc. 7300) has also been referred tc
in a

species.

this species.

Nearly

all

of the dorsal