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1†#&='!4<;+t+7Panax vietnamensis k@$Ÿ,~LQ16'C$C#$@0#$C
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.,4D#t+“JG71D#0#~$/1#'1%& #&?)u$$@'?#1;/$%1:•&'u#4/
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6¢: rw$"$#4/z1$C~1;/<;+&=1'#PLQ1#;#&'%#&$4#1A#&$|
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6'($,45q#5'#$C($C64D+N'#C1#$'C$
8 z ,4Drq#t#&6'($ ,4)q#6'($ 1Ž $C64D
'($ !45z6'C#9'1Ž $C64D$|$@v#&$u'$C64D)A'1DC#r'$?

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Ru1$C64D3…#&$@D#&+A'$@LM#&1B/1rq#t#&6'($ ,1B$@x1,4
1„1#t#&@D#&#s#&'r'(#$}1Q)~1$?!4+1D#s#&$C64D#4/$@•!v'
$@v#&$u'1Ž $C64D$'0#P5'#@ 1x#&8$C64D)A',4:u$@>#&='!4:u
$@>#)q#6'($ >,"/#s#&$C64D14#&&#,N'$@v#&$u'1Ž $C64D)A'6 D
#'0$>rq#t#&#A'1B/$4#1A#&14#&1 D6B/#'0
i8B6jEO
8 ;## #&'%#&1;/$@’#&
8 qD:q##&’#&y#
8 vD1;/5v16(#
8 <q#•B$1u1Dv$1B$5'#=1
8 T #&$}#1A#&#&'()
<\]^ E
 
XR@SURkU6Fl96P>>?=m8O
8 KKG•D6y@$DDry: #5u$LQ1$C64D5%#&3LN'r}#'?#,',4
L @ ru'#'(+$C64D
8 JEJT $$' 5<1!y'3y#,4yD3D@y<1˜ ##•LN#&=1$/C$
$C64D
8 “FI ##'&#A'1B/)A'$'0#•1u1!D4'=1q'
8 “EID&!•u1p#, '$@†1Ž 
8 “G<rDD&#&'0#1„5‚3-#&1u1Du1B$'D45'#$@L•#&,4
)u$5'#1n: #
8 “G*TD@y!,4T @$'#$z1'(#,'&”)invitro$4#1A#&
8 “K*T@ 5'&y,4<rDD&)u$+'#+A'$@LM#&#A'1B/$C64D$z1
,"$
8 “KI ,4T y5˜ @'!#$'0#$4#1A#&$@D#&$vDLQ11;/
n#62'$|#A'1B/6 D)B#1Ž 1;/14@%$
8 “K“;#!")$C64D$@#$|#A'1B/$C64D3p1!¢#&./#)…7
1Ž Hapopappus gracilis

8 “mEu$'(#R/$Dr'#'#1rq#t#&)u#&Ž•Gerberas
8 “mJT @$D#,412#&5z•;/3z#&:/$@>#1/?#&y#,4D$C64D
$@#
8 “JERA#&$/T'$5'y$@D1y+'1 !5!#$'0#P5q#•B$1B$$@ D
9'$„1B)$@0#:/+A1A#&#&'()6–#&#A'1B/$C64D3p1!¢#&'$D5)y@++
5))T'$5'Fy$@D1y+'1 !5
8 “JJ!y'#,412#&5z$u'5'#1;/1/?#&y#6–#&)Ln#&)u)6 #
&y#
8 ““ILn#&+v' &'%#&141 1/?#&y#¤! ,@< ,@
-)-PUI`>6Da>8D<=UVb
-)-)D<=UVb96<=
A')u$$@'?#:  '&' 'Dv#3p3L{#&,4$z3L{#&¥&' 'Dv#3p3L{#&
1#11u11B$'† 5'#$@L•#&?)u$$@'?#
@D#&#A'1B/)A'~LM#&#&, '$@†@B$: #$@=#&~LM#&51@D5y1D
#'rC$:q$%$n#&D4'@ +2$5%1B$$z#'0##L#LN13| ~3p11'C$+ !$~
<\]^ I
 
1 5y'#$Ž/);#!4#s#&1B$@B$1#$@D#&#A'1B/)A'%',N'1u11B$'†
5'#$@L•#&~ •'#$LM#&3…#&•#’#&2$B)~r'#y$'#1, '$@†•16'($1D5z
)u$$@'?#1Ž )A'
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#A'1B/1##'($2~u#5u#&$B)n#)A')u$$@'?#$z#'0#'($2*G¦*
D
R
$}1Q)1D5'#$@L•#&,4#q/++1Ž )A'Ru1)A'$„1B)@B$r5'#
$@L•#&PDv$2#&r'1x#&LQ1$u1@M',4#A'•'r'(#1'C5u#&IFFF
!•D•1n#$@D#&I&'M1'C5u#&5%$I#&4/#A'$'0#
w$"$#A'1B/)A'+ #&!v'#''(:q#L
8 ;#&'%#&1;/$@’#&
8 ‚5„15%#&1Ž )A'v$

8 /$@>)A'/C,41„)A'! '• 
8 <q#•B$v$#;#$vD+46q#1B$!4$C64D)A'
-)--D<=UVb;<H7UXnDE8>PU6?A=
@D#&#A'1B/+A,41n: #$u1@M'~r;1=#+O1$+: #$@=#&•1
6'($~+O)q'•$>#$@v#&5'#!•$%$,4 #&)u$$@'?#
13'#3L{#&r'#A'1B/1u162)"#ru1# 1Ž 1;/!4ru1# 
#L#&1+2$5%/011#&#L#&’#1 @6D#~#&’#LM#&~#&/0#$%
!LQ#&~,'!LQ#&,41u11B$'† 5'#$@L•#&T%#3/$@>5'#$@L•#&,4)u$
$@'?#1Ž 1n: ##A'1B/1#$LM#&•/0#1B/1/#: +A'$@LM#&+N'
%',N'#A'1B/+A1#695#&$0+1u11B$s1n1„ }$#'$n3LN'3v#&
1'3y +'#y~LM#&,4'#D5'$D!@D#&$@LM#&Q)#A'1B/+A~1u11B$'†
5'#$@L•#&1, '$@†: #$@=#&n#,>1u1+A$u1@M'rA#&1rq#t#&$9#&Q)
1u11B$#4/
A'1B/+A,41n: #$u1@M'LQ1„#&3-#&$@D#&
8 &'0#1„'r'(#5'#$@L•#&%',N'+2$62)"#D•1+2$+A1Ž 1;/
8 ;#1;/invitro
8 vD+A5•D)-1,-1D1u1#&'0#1„#L1=#3†#&$C64D~2$6'C#5D+ ~
o
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 
-)-,D<=UVb;<96o8F=86
•1'?+1Ž +A);#5'#!41„ 1u1$C64D#D#$@§~);#1' +v#~rA#&
6p,'@5•;+#")~,>,"/+A);#5'#!4+A3/#B$1Ž 1;/5v1,'@5TA);#
5'#$LM#&!41u1+A‡#1’',414#Ru1+A);#5'#3…#&?#A'1B/$LM#&
$u1$|1u1++#D#~1u11’'+N'>#$4#D•11u114##D#
@D#&#A'1B/5z1;#6–#&&'s 1u11B$'† 5'#$@L•#&@B$: #$@=#&
T%#$vD1’'1#695#&1/$Dr'#'#D•1$9Q)1/$Dr'#'#,N' •'#~+%#$vD@ˆ
$>695#& •'#
A'1B/+A);#5'#LQ15‚3-#&$@D#&
8 vD1;/5v1,'@5

8 ;#&'%#&invitro
8 vD1;/ 62': •‚!•1D!1'1'#
8 &'0#1„:u$@>#>#$4#1n: #
-)-0D<=UVbdEJ96V8H76p>96V8
W'?+1Ž #A'1B/6 D)B#!4n#&'q#,$ D$u1rw$"$,4+A'$@LM#&
#A'1B/~#L#&1$?$vD@ 1q1;/!L{#&62'$|5D+ 1Ž $4#6 D)B#~3D,"/
5¨r);#6'($,N'1;/$z!L{#&62'$|1;/n#62'
v$)B##A'1B/1$?)u$$@'?#$4#1;/n#62'D4#1‡#$@D#&'
r'(##A'1B/invitro6–#&1D#LM#&$vD)A'$@z1$'C)D•1&'u#$'C)$A#&: $vD
+A5•D,4$vD1n: #Ln#&)u)$vD1;/n#62'r”),4)Ln#&)u)1=#!=1
$vD&'%#&1'(:q1=#!=1@B$1 D~•16'($#CLQ1rC$Q),N'1u1)Ln#&
)u)$vD@ 1u16'C#3p3'$@/#ru1# #L! 's$}#D•1&;/2$6'C##;#
$vD
4#)#+A'$@LM#&#A'1B/$ /9'$…/$21,4Dr'?&y#,4$9'1Ž
6 D)B#1ƒ#&#L1u1'r'(#+4•1;/1D6 D)B#5'#$@L•#&‘u$@>#
5'#$@L•#&,5 1Ž )A'v$)B#†'¢'695#&1u1+%'rDu#&,4D+A'
$@LM#&#5%)u$5'#1;/n#62',45'#$@L•#&1Ž 1;/#'1#&5¨$%$n##C
<\]^ K
 
1x#&LQ1#A'$@D#&'r'(#1'C5u#&~+•13…1;/v$)B#1Ž +2$5%r'?
&y#5'#$@L•#&$@D#&1q ''r'(#11'C5u#&,4$@D#&$%'
A'1B/6 D)B#,4v$)B#LQ13…#&1D$vD1u13†#&$#?
8 &'0#1„&y#!•#,>1x#&rA#&6'?'(#•1n$?3pQ)$‚
8 R=#1u13†#&2$6'C#
-)-qD<=UVb>cd7JQX8
C64Dn#1$?#"#LQ16–#&1D#LM#&#&'#+A~D•1•‚!•y#Ÿ/+y
< 1x#&LQ1#A'1B/3p1!¢#&~1rB/D•1! 1$vD'r'(#$"#!Q'1D
5z$@ D9'r},4$'C)•x1,N'+A'$@LM#&3'#3L{#&C64Dn#LQ1!=1,4#A'
1B/$@D#&+A'$@LM#&•16'($,4$t#&5'#r%'
T¡'!Dv'1;/~+¡'!Dv'$C64Dru1# †'¢'#s#&rw$"$#A'1B/ru1

# \013'#3L{#&1D#A'1B/$C64Dn#ru)„1$v)~3D1x#&6p+B$
#'1B$1#$'C$1D5'#$@L•#&r'$u1@M'r¢':#$?$C64D
A'1B/$C64Dn#LQ15‚3-#&1D1u1+-1}1
8 R=#3†#&$C64D
8 &'0#1„1B$@x1$C64D~)u$$@'?#,4);# $C64D$@D#&#s#&'
r'(#ru1# 
8 #"#1u11B$$@ D9'$„1B)
-)-MD<=UVb9?J>J9REF>
C64D$@#!4$C64DPLQ1$u16¢$4#$C64D6–#&)Ln#&)u)1n=1
 /5‚3-#&y#Ÿ/+y@D#&'r'(##A'1B/)…Q))@D$D)! 5$1$?$u'5'#
$4#$C64D+N'~);#1' ,4$u'5'#$4#1;/D4#1‡#W$C1Ž rw$"$
$u1,4#A'1B/$C64D$@#!4$C64DrA#&1+4#&1„#&~•$@v#&$u'n#64D~+"$
2$C64D$LQ1$@0#n#,p$?$}1+A'$@LM#&1$?@B$1 D.v$F
K
$C64DH+!
+A'$@LM#&7
©#&3-#&#A'1B/)@D$D) 5$
8 vD@ 1;/! '5D+ 
8 R/?#1u164D: #,41/?#&y#‘u$@>#3#&Q))@D$D)! 5$1$?
LQ1$z1'(#$@0# '%'$LQ#&1…#&!D4' /ru1!D4'
<\]^ m
 
8 ‘u$@>#5'#$9#&Q)+4#&$C64D
-),<=>?@A8B8D<=UVb
4#)#+A'$@LM#&#A'1B/&’+#t+$4#)#1n6q#5 
8 T%'rDu#& !LQ#&
8 T%'rDu#&,'!LQ#&
8 '$ +'#
8 &’#1 @6D#
8 RB$'† 5'#$@L•#&

8 &D4'@ 1$?695#&1u1$4#)#rA#&•u1p#.#LN13| ~3p11'C$
#B++y#~o7,4 & @
TA'$@LM#&#A'1B/1$?1' $4#6 !Dv'
8 TA'$@LM#&#&•D3'#3L{#&ª'$y~#D)
8 TA'$@LM#&14+!LQ#&1B$3'#3L{#&$@#&6># +6D@&
8 TA'$@LM#&&'43'#3L{#&T<
Rn5•1D,'(1•;/3z#&1u1+A'$@LM#&#A'1B/!4•y+•”$1u1$4#)#
1#1D5z5'#$@L•#&,4)u$$@'?#1Ž 1;/
4#)#+A'$@LM#&#A'1B/$C64D,4+A$z1,"$$ /9'$…/$yD!D4'
,462)"##A'1B/%',N'1…#&+2$+O1B/#L#&$…/$yD+-1}1$}#&'(+
$>$4#)#+A'$@LM#&1ƒ#&5¨$ /9'$…/$yD&' 'Dv#);# 1Ž +O1B/
C64D+A$z1,"$†'¢')$%'L1D5'#$@L•#&,4)u$$@'?#$@D#&#A'
1B/2)1Ž +A'$@LM#&3'#3L{#&q#L•#&$@z1$'C)$N':u$@>#$#"#
1u11B$3'#3L{#&$|+A'$@LM#&,4D$C64D>,"/1#'1‡#2)+A'
$@LM#&$}1Q)$@LN1r'#A'1B/LM#&5‚3-#& ` /R!!DP#&?'
1‡#2)
<\]^ J
 
-)0PUU6V>Q=ZD6rEF=86>?@s8B>6eUHa>
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8 4+!LQ#& •'##2'5'#1Ž +O1B/
8 q#t#&$9#&Q) •'#$z#'0#1Ž +O1B/

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8 }1$}15z);#Du1Ž 1u1+A3O#
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8 4+$t#&3'(#$}1)'C#!u3Dr}1$}15z!N#!0#1Ž $C64D
8 R$?!4+$t#&5z+•1Ž r}r9#&•+2$5%!D4'
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8 u#&Žv$&'%#&D•11Ž&'%#&
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8 R$?&;/$vD:qrA#&v$D•1!4+$t#&r}1$LN1:q#DrA#&v$
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8 R$?!4+1"+5zDu&'4•!u,4:q1;/1+x'
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t+“KJ~<ƒ„1T'#P$>+$B/1;/<;+&=1'#$v',…#&#x'
&=1'#,4$v+•$$0##!4<;+Gt+“mE~D4#'$@ 1;/$%11Ž
 #3;#/G3D<4D'+D#&3O#P)u$'(#+2$!D4'Panax +=1
D #&$4#:#$?•21 DJFF+$@0#,…#&#x'&=1'#,4$v+•$$0#!4
<;+%$@x1,N'$0#rD =1 Panax articulatus, = ;#  <;+  .Araliaceae7
.&/ˆ# T'# „1~  *FFE7  0#  rD  =1 1Ž  1;/  LQ1  1A#&  #"#  !4 Panax
vietnamensis  y$@5~=;#<;+@ !' 1y y~1A#&6%$v''(#$z1,"$
 + @D,.'0#eA$@LN1;/7#°+“JG3D4p#&,4@5,'5$r/•$
$0#.'#$y@#y$I7
•86-)Oo;ByU=868BJ7=>e86=g8
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