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giáo trình giảng dạy trực tuyến
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:HEVLWHKWWSZZZFWXHGXYQNQQHPDLOGPLQK#FWXHGXYQYWDQK#FWXHGXYQ






Giáo trình
dinh dỡng gia súc

chơng 4
tiêu hóa và hấp thu




Chủ biên: PTS. Lu Hữu Mãnh
Tham gia biên soạn: Th.S. Nguyễn Nhựt Xuân Dung
và PTS Võ Văn Sơn

Tháng 1 năm 1999



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ir1+1*+p$
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QKQJSKQWQJLDQ[D\UDEQWURQJQJWLXKRDFXDQJYWQKQJSKQ
WQD\FRWKFKSWKX&DFTXDWUQKWLXKRDTXDQWURQJFRWKQKRPODL
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 $6m7,j8+2v$

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QJFXDVRQJQKXQJGRVFRERSFDFFFXDYDFKWEDRUXW&KFQQJ
FXDWDFQJFJLLQ\ODYQFKX\QWKFQWUQWKFQYLGLFKWLXKRD
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 7KFQFWLXKRDGDGD\WQJSKQFD[XQJUXWQRQQL\
WKFQVHFWDFGXQJYLFDFGFKFXDWDWUDQJJDQYDWX\'FKWUDQJWLW
UDPWPLWUQJEDVHEDRYYDFKWDWUDQJGRV[PQKSFXDNKLWKF
QWPDFLGFKORUK\GULF
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 0WGRJDQEDLWLWUDFKDFDFPXLQDWULXPYDNDOLXPFXDFDFDFLGPWFKX
\X OD FDF JO\FRFKROLF YD WDXURFKROLF ELOLYHUGLQ YD ELOLUXELQ OD VF W PW
QJRDLUDFRQFRFKROHVWHUROYDPXFLQ&DFPXLPWRQJYDLWURTXDQWURQJ
TXDWUQKWLXKRDEQJFDFKKRDWKRDHQ]\PHOLSDVHYDQKXWQJKRDFKW
EHR
 'FKWX\GRWX\WDQJWLWUDFRPWV\XWWX\WDQJEDLWLWUDGFKWX\

NKLDFLGDYDRWDWUDQJKRUPRQVHFUHWLQFJLDLSKRQJUDWOSELXE
FXD UXWQRQDYDR WURQJ PDX.KL QR WLS[XFYL VW XQ KRDQ FXD WX\
WDQJ QR NFK WKFK FDF W EDR WX\Q WX\ WLW UD PW FKW GFK FKD FDF LRQ
ELFDUERQDWH YL QQJ  FDR QKQJ UW W HQ]\PH 0W KRUPRQ NKDF
FKROHF\WRNLQLQSDQFUHR]\PLQFXQJFPDQJQK\WLWUDNKLFDFSHSWLGYD
NKL WKF Q WLX KRD WLQ WL WD WUDQJ +RUPRQ Q\ NFK WKFK EDL WLW FDF
HQ]\PH QK WU\SVLQRJHQ FK\PRWU\SVLQRJHQ SURFDUER[\SHSWLGDVH $ YD %
SURODFWDVH αDP\ODVH OLSDVH OHFLWKLQDVH YD QXFOHDVH .KQJ JLQJ QK
SHSVLQFDFHQ]\PQ\KRDWQJS+W(QWHURNLQDVHVODPWHQ]\PH
GR PDQJ QK\ FXD WDWUDQJFKX\QLWU\SVLQRJHQNKQJKRDWQJWKDQK
WU\SVLQ+RDWQJQ\FXQJF[XFWDFGRFKQKELWU\SVLQYDQKWKWLS
WXFFDFSKDQQJ[XFWDFWQJKRD+RDWQJFXDWU\SVLQUWFELWQR
FKWDFGXQJOQOLQNWSHSWLGEDRJPQKRPFDUER[\OFXDO\VLQYDDUJLQLQ
7U\SVLQFXQJFKX\QLFK\PRWU\SVLQRJHQWKDQKHQ]\PHFK\PRWU\SVLQF
ELW WDF GXQJ OQ OLQ NW VQ SHSWLG EDR JP QKRP FDUER[\O FXD W\URVLQ
WU\SWRSKDQSKHQ\ODODQLQYDOHXFLQ
 (Q]\PH αDP\ODVH FXD GFK WX\ FR FKF QQJ WQJWDP\ODVH FXDWX\Q
QFERWFRWDFGXQJOQOLQNWαJOXFDQWLQKEWYDJO\FRJHQ
 6SKQJLDLFKWEHRFKX\XGRWDFGXQJFXDOLSDVHGFKWX\(Q]\PHQ\
NKQJ WKX\SKQKRDQWRDQFDFWULDF\OJO\FHUROYDKRDWQJGQJODLJLDL
RDQPRQRDF\OJO\FHURO
 &KWEHRULGDGD\GDQJSKQWOQUWNKREWKX\SKQQKDQKVWKX\
SKQFKWEHRFKWUELVQKXWQJKRDFXDPXLPW
 6WLXKRDUXWJLD
 9LQKQJNKXSKQEQKWKQJOXQOXQFRPWOQJWKFQNKDQJ
WDF GXQJ FXD FDF HQ]\PH WLX KRD lt KHR FHOOXORVH  YD FDF KHPLFHOOXORVH
NKQJ EFDFHQ]\PHWURQJEPD\WLXKRDWDFGXQJ/LJQLQODWKDQKSKQ
NKQJWLXKRDYKRDQWRDQNKQJEFDFHQ]\PWLXKRDWDFQJ



 &QJKRDWQJFXDYLVLQKYWFKX\X[D\UDPDQKWUDQJFDHFXP
&DFYLNKXQKLQGLQJPFRODFWREDFLOOLVWUHSWRFRFFLFROLIRUPEDFWHURLGHV
FORVWULGLDYDQPPHQ1KQJYLVLQKYWQ\WUDRLWUQNKXSKQYDFDF
FKWEDLWKDLQLVLQKKXTXDODUXWJLDKQKWKDQKFDFWKLQGROHVNDWROH
SKHQROK\GURJHQVXOSKLGHDPLGDPPRQLDYDFDFDFLGEHRED\KLQKQK
DFLGDFHWLFSURSLRQLFYDEXW\ULF7UQKHRPWVQJKLQFXYPFWLXKRD
FKRWK\UQJFKW[FXDPWVORDLWKFQESKQJLDLWURQJPWSKDPYL
JLLKDQ6SKQJLDLGRYLVLQKYWFKX\X[D\UDUXWJD6WLXKRD
FHOOXORVHYDFDFSRO\VDFFKDULGFDRSKQWKHRUWWKSVRVDQKYLQJDYD
WKXQKDLODLNKDF
 9LNKXQFXDUXWJLDFRQWQJKSFDFYLWDPLQQKRP%VHFYWFKXKS
WKXFKRFWK7X\QKLQVWQJKSFDFQKRPYLWDPLQ%WEPD\WLXKRD
OD NKQJ X  DS QJ QKXFXKDQJQJD\YY\FQSKDLFXQJFSFDF
YLWDPLQQ\WWKFQ
  7RPODLVWLXKRDWKXFYQKVDX
DlPLQJWLQKEW
3W\DO LQ
GH[WULQ
EltGDGD\3URWHLQ
SHSVLQ
SRO\SHSWLGSHSWRQГWDFLGDPLQ
  F2tWDWUDQJFRGFKWX\
 3RO\SHSWLGSHSWRQ
7U\SVLQ
DFLGDPLQWSHSWLGSHSWRQ
   7LQKEWαDP\ODVHPDOWRVHJOXRVH
   /LSLG 
/LSD]QLPPD£F''
PRQRJO\FHULG


  GltUXWQRQFRGFKWUDQJ
     1+
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SHSWLGDVH  GLSHSWLGDVH
  3RO\SHSWLGSHSWRQ    SHSWLGDD   DD

   0DOWRVH  0DOWDVH  *OXFRVH   
/DFWDVH
 /DFWRVH    JOXFRVHГJDODFWRVH
