Phân lập và xác định cấu trúc hợp chất steroid từ lá thân cây gai (boehmeria nivea (l) gaud ) họ gai urticaceae

751*,+&+ễ1*91*
.+2$.+2$+&71+,ầ1
----------------------------------

1*é7+7+ễ</,1+

3+ặ1/39ơ;ẩ&1+&875ề&
+3&+767(52,'7/ẩ-THN CY GAI
(Boehmeria nivea (L) Gaud
+Ӑ*$,Urticaceae)

.+Ï$/8Ұ17Ӕ71*+,ӊ3ĈҤ,+Ӑ&
Ngành 6ѭSKҥP+yDKӑF

3K~7Kӑ


751*,+&+ễ1*91*
.+2$.+2$+&71+,ầ1
----------------------------------

1*é7+7+ễ</,1+

3+ặ1/39ơ;ẩ&1+&875ề&
+3&+767(52,'7/ẩ-THN CY GAI
(Boehmeria nivea (L) Gaud
+Ӑ*$,Urticaceae)

.+Ï$/8Ұ17Ӕ71*+,ӊ3ĈҤ,+Ӑ&
1JjQK6ѭSKҥP+yDKӑF

*,Ҧ1*9,Ç1 +ѬӞ1*'Ү17K61JX\ӉQ7Kӏ1JD
+ӑFYLӋQSKzQJNK{QJ-khơng qn
2. 767ULӋX4Xê+QJ
7UѭӡQJĈҥLKӑF+QJ9ѭѫQJ

Phú 7Kӑ18


/Ӡ,&$0Ĉ2$1

7{L[LQ FDP ÿRDQÿk\ Oj F{QJWUuQK QJKLrQ FӭX FӫD ULrQJW{LGѭӟLVӵ
KѭӟQJ GүQ NKRD KӑF FӫD ThS. 1JX\ӉQ 7Kӏ 1JD và 76 7ULӋX 4Xê +QJ.
&iF Vӕ OLӋX NӃW TXҧ QrX WURQJ .KyD OXұQ Oj WUXQJ WKӵF Yj FKѭD WӯQJ Fy DL
F{QJ Eӕ WURQJ FiF F{QJ WUuQK QJKLrQ FӭX NKRD KӑF WUѭӟF ÿk\ 7RjQ Eӝ FiF
WK{QJWLQWUtFKGүQWURQJ.KyDOXұQÿmÿѭӧFJKLU}QJXӗQJӕF[XҩW[ӭ

Sinh viên

1J{7Kӏ7K\/LQK


/Ӡ,&Ҧ0Ѫ1
.KyDOXұQQj\ÿѭӧFWKӵFKLӋQGѭӟLVӵKѭӟQJGүQFKӍEҧRWұQWuQKFӫD
ThS. 1JX\ӉQ7Kӏ1JDvà 767ULӋX4Xê+QJ9ӟLOzQJNtQKWUӑQJYjELӃW
ѫQVkXVҳFHP[LQFKkQWKjQKFҧPѫQWKҫ\F{ÿmWұQWuQKFKӍEҧRNKtFKOӋ
KѭӟQJGүQÿӝQJYLrQYjJL~SÿӥHPKRjQWKjQKNKyDOXұQWӕWQJKLӋS
EP[LQFKkQWKjQKFҧPѫQTXêWKҫ\F{BӝP{Q+yDKӑFFQJFiFWKҫ\
cô .KRD.KRDKӑF7ӵQKLrQ-TUѭӡQJĈҥLKӑF+QJ9ѭѫQJÿmWұQWuQKJL~S
ÿӥWҥRPӑLÿLӅXNLӋQWKXұQOӧLYӅWKӡLJLDQYjWjLOLӋXgiúp HPKRjQWKjQKÿӅ
tài NKyDOXұQWӕWQJKLӋS

1KkQ GӏS Qj\ HP [LQ Ej\ Wӓ Vӵ FҧP ѫQ VkX VҳF ÿӃQ %DQ /mQK ÿҥR
7UѭӡQJ ĈҥL KӑF +QJ 9ѭѫQJ .KRD .KRD KӑF 7ӵ QKLrQ-7UѭӡQJ ĈҥL KӑF
+QJ 9ѭѫQJ ÿm JL~S ÿӥ WҥR FKR HP PӝW P{L WUѭӡQJ KӑF WұS NKRD KӑF
nghiêm túc.
&XӕLFQJHP[LQEj\WӓOzQJELӃWѫQVkXVҳFÿӃQJLDÿuQKQJѭӡLWKkQYj
FiFEҥQVLQKYLrQOӟSK12 Ĉ+63+yDKӑF-7UѭӡQJĈҥLKӑF+QJ9ѭѫQJ và
ÿһF ELӋW FiF EҥQ WURQJ QKyP NKyD OXұQ WӕW QJKLӋS ÿm Kӛ WUӧ ÿӝQJ YLrQ HP
WURQJVXӕWTXiWUuQKKӑFWұSYjWKӵFKLӋQNKyDOXұQ.
Em xLQFKkQWKjQKFҧPѫQ WҩWFҧVӵJL~SÿӥTXêEiXQj\!
3K~7K͕QJj\ 02 tháng 5 QăP8
Sinh viên

1J{7Kӏ7K\/LQK


-i-

0Ө&/Ө&

'$1+0Ө&&È&.é+,ӊ8&+Ӳ9,ӂ77Ҳ7 ............................................ iii
'$1+0Ө&&È&+Ỵ1+ ................................................................................ iv
'$1+0Ө&&È&%Ҧ1*................................................................................ v
0ӢĈҪ8 ........................................................................................................... 1
/tGROӵDFKӑQÿӅWjL...................................................................................... 1
0өFWLrXÿӅWjL ............................................................................................... 2
éQJKƭD NKRDKӑFYjWKӵFWLӉQFӫDÿӅWjL ...................................................... 2
&+ѬѪ1*7Ә1*48$17¬,/,ӊ8 ........................................................... 4
*LӟLWKLӋXYӅFKLBoehmeria ....................................................................... 4
ĈһFÿLӇPWKӵFYұWFk\*DL .......................................................................... 9
&iFQJKLrQFӭXYӅKyDWKӵFYұWFKLBoehmeria........................................ 10

&+1*,71*97/,89ơ3+1*3+ẩ31*+,ầ1
&8 ................................................................................................................ 15
LWQJQJKLrQFX................................................................................ 15
9WOLXWKLWEQJKLrQFX ...................................................................... 15
+yDFKW.................................................................................................. 15
7KLWEQJKLrQFX ................................................................................ 15
3KQJSKiSQJKLrQFX ........................................................................... 16
3KѭѫQJSKiSQJkPFKLӃW ........................................................................ 16
3KѭѫQJSKiSVҳFNê ............................................................................... 16
2.3.3KѭѫQJSKiSVҳFNêOӟSPӓQJ ............................................................ 17
3KѭѫQJSKiSVҳFNêFӝW ...................................................................... 18
3KѭѫQJSKiSNӃWWLQK ............................................................................. 19
3KѭѫQJSKiSSKәFӝQJKѭӣQJWӯKҥWQKkQ ............................................ 19
3KәFӝQJKѭӣQJWӯQKkQSURWRQ1H-NMR) ...................................... 20


-ii-

3KFQJKQJWQKkQFDUERQ-13 (13C-NMR)................................. 21
&+1*.7481*+,ầ1&89ơ7+2/81 ........................ 23
3KkQOSFQGFKFKLWn-hexane lỏ-thõn cõy Gai .................................... 23
LXFKFQFKLWQ-hexane .................................................................. 23
4XiWUuQKSKkQOұSKӧSFKҩWVWHURLG ....................................................... 25
.KҧRViWWKjQKSKҫQÿӏQKWtQKYjOӵDFKӑQGXQJP{L......................... 25
4XiWUuQKSKkQOұSFKҩW ....................................................................... 27
;iFÿӏQKFҩXWU~FKӧSFKҩW%1& ............................................................ 33
.ӂ7/8Ұ19¬.,ӂ11*+ӎ......................................................................... 38
.ӃW OXұQ ....................................................................................................... 38
.LӃQQJKӏ ..................................................................................................... 38
7¬,/,ӊ87+$0.+Ҧ2 ............................................................................... 39

-iii-

DANH MӨC CÁC KÝ HIӊU, CHӲ VIӂT TҲT

™ &iFSKѭѫQJSKiSVҳFNê
TLC

7KLQ/D\HU&KURPDWRJUDSK\6ҳFNêOӟSPӓQJ

CC

&ROXPQ&KURPDWRJUDSK\6ҳFNêFӝW

™ &iFSKѭѫQJSKiSSKә
1

H-NMR

Proton 1XFOHDU 0DJQHWLF 5HVRQDQFH 6SHFWURVFRS\ 3Kә FӝQJ
KѭӣQJWӯKҥWQKkQSURWRQ

s: singlet

d: doublet

t: triplet

m: multiplet

™ &iFFKӳYLӃWWҳWNKiF
CTPT

&{QJWKӭFSKkQWӱ

™ 7rQFӫDFiFKӧSFKҩWÿѭӧFYLӃWWKHRQJX\rQEҧQ7LӃQJ$QK


-iv-

DANH MӨC CÁC HÌNH
Hình 1.1. Boehmeria diffusa Wedd. ± 1JӑQFk\PDQJKRD ............................. 4
Hình 1.2. Boehmeria macrophylla Hornem. ± 1JӑQFk\PDQJKRD................ 5
Hình 1.3. B. nivea (L.) Gaud. ± 1JӑQFjQKPDQJKRD+RDFiL+RD
ÿӵF4Xҧ ........................................................................................................ 7
Hình 1.4. B. penduliflora Wedd. ± 1JӑQFjQKPDQJKRD................................. 8
Hình 1.5. B. tonkinensis Gagnep.±1. Cành mang hoa; 2. Xim co cái; 3. Hoa
ÿӵFQKuQӣWUrQ%ҫXKRD/iÿjLYjEDRSKҩQ .......................................... 9
Hình 1.6. Cây Gai ............................................................................................ 10
+uQK0LQKKӑDVҳFNêOӟSPӓQJ .............................................................. 17
+uQK0LQKKӑDVҳFNêFӝW ........................................................................ 18
+uQK0LQKKӑDSKә1H-NMR................................................................... 21
+uQK0LQKKӑDSKә13C-NMR.................................................................. 22
+uQK7KkQYjOiFk\*DLÿѭӧFWKiLQKӓSKѫLWURQJEyQJPiW ................ 23
+uQK4XiWUuQKQJkPFKLӃWFk\*DLÿLӅXFKӃFһQFKLӃWQ-hexane ................... 24
+uQK.ӃWTXҧNKҧRViW7/&FһQFKLӃWQ-Hexan FӫDFk\*DL ................... 26
+uQK&ӝWWәQJVLOLFDJHOFһQQ-hexane ..................................................... 28
+uQK.ӃWTXҧNKҧRViW7/&FiFEuQKKӭQJGXQJGӏFKUӱDJLҧL ............... 29
+uQK+uQKҧQK7/&FiFSKkQÿRҥQ)·) ............................................ 31

+uQK+uQKҧQK7/&FKҩW%1& .............................................................. 31
+uQK6ѫÿӗSKkQOұSFһQQ-hexane lá-thân cây Gai ................................. 32
Hình 3.97/&VRViQK%1&YӟLKӛQKӧSE-Sitosterol+Stigmasterol .......... 33
Hình 3.103Kә1H-105FӫDKӧSFKҩW%1& ................................................ 34
Hình 3.113Kә1H-105JLmQUӝQJ (1) FӫDBNC4 ......................................... 35
Hình 3.133Kә1H-105JLmQUӝQJ (2) FӫDBNC4 ......................................... 36


-v-

DANH MӨC CÁC BҦNG
%ҧQJ*LiWUӏ5f YjPjXVҳFFiFYӋWFKҩWWUrQEҧQPӓQJ .......................... 27
%ҧQJ.ӃWTXҧFiFSKkQÿRҥQWKXÿѭӧFWӯFӝWWәQJQ-hexane .................. 30


-1-

MӢ ĈҪU
1. Lí do lӵa chӑQÿӅ tài
7ӯ [D [ѭD FRQ QJѭӡL ÿm ELӃW Vӱ GөQJ FiF ORjL Fk\ Fӓ Fy VҹQ WURQJ Wӵ
QKLrQÿһFELӋWOjFiFORjLFk\FӓÿmFyWURQJUӯQJÿӇOjPWKXӕFFKӳDEӋQKWӯ
FiF ORҥL EӋQK WK{QJ WKѭӡQJ FKR ÿӃQ FiF ORҥL EӋQK NKy FKӳD WUӏ 7KHR WKӡL
JLDQFQJYӟLVӵSKiWWULӇQNKRDKӑFQyLFKXQJYjFӫDQJKjQK\KӑFQyLULrQJ
con QJѭӡLÿmELӃWVӱGөQJFiFKӧSFKҩWWKLrQQKLrQWәQJKӧSQKLӅXORҥLKӧS
FKҩW KӳX Fѫ SKөF Yө cho QKX FҫX OjP WKXӕF FKӳD EӋQK QkQJ FDR VӭF NKӓH
FӝQJÿӗQJ. 1JXӗQVLQKYұWÿһFELӋWOjFiFFk\WKXӕFFәWUX\ӅQYүQWLӃSWөFOj
PӕLTXDQWkPÿһFELӋWFӫDFiFQKjQJKLrQFӭXYӅFiFKӧSFKҩWWKLrQQKLrQFy
KRҥWWtQKVLQKKӑFYuWtQKÿDGҥQJFӫDFK~QJWKѭӡQJKӭDKҽQVӵSKRQJSK~
FӫDFiFFKҩWKyDKӑFYӟLFҩXWU~FPӟLOҥ
+LӋQQD\FiFKӧSFKҩWFyQJXӗQJӕFWKLrQQKLrQÿyQJJySVӕOѭӧQJ

WKXӕFWUrQWKӏWUѭӡQJWKӃJLӟL YjÿyFNJQJOjWӹOӋÿӕLYӟLFiFKӧSFKҩWÿDQJ
WURQJJLDLÿRҥQWKӱQJKLӋPOkPVjQJ7KHRWKӕQJNrWKutWQKҩWFyKӧS
FKҩWEҳWQJXӗQWӯORjLFk\FyWKӇÿѭӧF[HPOjFiFWKXӕFTXDQWUӑQJÿѭӧF
Vӱ GөQJ ӣ PӝW KD\ QKLӅX QѭӟF &k\ WKXӕF GkQ JLDQ Wӯ OkX ÿm ÿѭӧF QKLӅX
QJѭӡLTXDQWkPÿӃQÿk\OjQJXӗQWjLQJX\rQWKӵFYұWFyJLiWUӏWKLӃWWKӵFFKR
FiFFӝQJÿӗQJÿӏDSKѭѫQJWURQJYLӋFSKzQJFKӳDEӋQKQJRjLUDQyFzQFyJLi
WUӏEҧRWӗQQJXӗQJHQFXQJFҩSFKROƭQKYӵFGѭӧFKӑF9LӋW1DPOjQѭӟFFy
ÿӏDKuQKYjNKtKұXWKXұQOӧLFKRFiFORjLWKӵFYұWSKiWWULӇQ EұFQKҩWĈ{QJ
1DPÈOjQѫLWұSWUXQJQKLӅXFk\WKXӕFTXêKLӃPYӟLGkQWӝFVLQKVӕQJYj
KӑFyWUX\ӅQWKӕQJOkXÿӡLWURQJYLӋFVӱGөQJQJXӗQWjLQJX\rQWKӵFYұWNgày
nay P{LWUѭӡQJÿDQJFyQJX\FѫEӏ{QKLӉP FRQQJѭӡLFNJQJSKҧLÿӕLPһWYӟL
QKLӅX QJX\ Fѫ EӋQKWұWNKiF QKDX Yu Yұ\ YLӋFVӱ GөQJ FiF ORjL WKXӕFWURQJ
WKLrQQKLrQQJj\FjQJÿѭӧFѭDFKXӝQJ


-2-

9ӟLKӋWKӵFYұWSKRQJSK~YjÿDGҥQJQKѭYұ\FK~QJWDSKҧLWұQ GөQJ
WӕLѭXFiFQJXӗQWKӵFYұW. +LӋQQD\FiFF{QJWUuQKQJKLrQFӭXWuPUDFiFKӧS
FKҩW PӟL FNJQJ QJj\ FjQJ SKiW WULӇQ ÿӇ Fy WKӇ NKiP SKi WLӃS FұQ Yj WuP UD
QKLӅXKѫQQӳDF{QJGөQJFӫDFiF FKҩWFӫDFiFORҥLFk\WURQJ\KӑFPӝWFiFK
QJKLrPW~FFKLWLӃWYjrõ ràng. &QJYӟLVӵSKiWWULӇQFӫDF{QJQJKӋWәQJKӧS
KyDGѭӧFWҥRUDFiFELӋWGѭӧFFiFQKjNKRDKӑFYүQÿDQJFӕJҳQJWuPKLӇX
NKiPSKiFiFKRҥWWtQKVLQKKӑFNKiFQKDXFӫDFiFKӧSFKҩWFyQJXӗQJӕFWӯ
QKLӅXORjLWKӵFYұWNKiFQKDX
Tên chi Boehmeria ÿѭӧFÿһWWKHRWrQQKjWKӵFYұWKӑFQJѭӡLĈӭF Georg
Rudolf Boehmer. Theo FXӕQ ³7ӯ ÿLӇQWKӵF YұW WK{QJGөQJ´FӫD WiFJLҧ 9}
9ăQ&KLFKLBoehmeria FyNKRҧQJORjLӣQKLӋWÿӟLYj%ҳFFұQQKLӋWÿӟLӢ
QѭӟFWDFyORjLPjORjLÿmELӃWF{QJGөQJ ÿѭӧFVӱGөQJWURQJ\KӑFFә
WUX\ӅQ[3] PӣUDWLӅPQăQJQJKLrQFӭXKyDWKӵFYұWYӅFiFORjLQj\

7LӃS WөF SKiW WULӇQ KѭӟQJ QJKLrQ FӭX YӅ WKjQK SKҫQ KyD KӑF FiF ORjL
WKӵF YұW WUrQ ÿӏD EjQ WӍQK 3K~ 7Kӑ QKҵP PөF WLrX SKiW KLӋQ FiF KӧS FKҩW
WKLrQQKLrQWӯFiFORjLWKӵFYұWFӫD9LӋWNam, tôi OӵDFKӑQÿӅWjL³3KkQO̵S
Yj[iFÿ͓QKF̭XWU~FKͫSFK̭WsWHURLGWͳlá- thân cây Gai (Boehmeria nivea
(L) Gaud
K͕*DL8UWLFDFHDH´.
2. MөFWLrXÿӅ tài
- 3KkQOұSÿѭӧFKӧSFKҩWVWHURLGWӯFһQGӏFKFKLӃWn-KH[DQHFӫDOi- thân
cây Gai.
- ;iF ÿӏQK FҩX WU~F KӧS FKҩW VWHURLG SKkQ OұS ÿѭӧF EҵQJ FiF SKѭѫQJ
SKiSSKәFӝQJKѭӣQJWӯKҥWQKkQ
3. éQJKƭDNKRDKӑc và thӵc tiӉn cӫDÿӅ tài
- éQJKƭDNKRDK͕F
+ .ӃWTXҧFӫDÿӅWjLÿyQJJySWK{QJWLQFKROƭQKYӵFQJKLrQFӭXKyDKӑF
FiFKӧSFKҩWWKLrQQKLrQ YӅWKjQKSKҫQKyDKӑFFӫD cây Gai (Boehmeria nivea
(L) Gaud.) Kӑ*DL (Urticaceae) FӫD9LӋW1DP


-3-

+ &XQJ FҩS FiF Gӳ OLӋX YӅ SKә FӝQJ KѭӣQJ Wӯ KҥW QKkQ FӫD KӧS FKҩW
steroid WӯFk\*DL
&XQJFҩSWK{QJWLQYӅFiFNƭWKXұWQJkPFKLӃWVҳFNt VҳFNtEҧQPӓQJ
YjVҳFNtFӝW
[iFÿӏQKFҩXWU~FFӫDFiFKӧSFKҩWWKLrQQKLrQ
- éQJKƭDWK͹FWL͍Q
ĈӅWjLJiúp ÿӏQKKѭӟQJFKR YLӋFQJKLrQFӭXVӱGөQJFk\*DL trong
FXӝFVӕQJÿӗQJWKӡLJySSKҫQQkQJFDRNLӃQWKӭFYӅSKkQOұS YӅSKәFӝQJ
KѭӣQJWӯKҥW nhân cho sinh viên.
ĈӅWjLFzQOjPWjLOLӋXWKDPNKҧRFKRVLQKYLrQQJjQKKyDKӑFYjFiQ

EӝQJKLrQFӭXKyDKӑFKӳXFѫ


-4-

&+ѬѪ1*. TӘNG QUAN TÀI LIӊU
1.1 Giӟi thiӋu vӅ chi Boehmeria
Theo WjLOLӋX´ 7ͳÿL͋QWK͹FY̵WWK{QJGͭQg ³FӫDWiFJLҧ9}9ăQ&KL[1],
Boehmeria -DFTKӑ*DL ± 8ULFDFHDH>ÿӇWѭӣQJQKӟWKҫ\WKXӕFYjQKjWKӵF
YұW KӑF ĈӭF *HRUJ 5XGROI %RHKPHU -1803)] ± tên TLӃQJ VLӋW Oj Gai,
*DLWX\ӃW
+uQKGҥQJOiPӑFÿӕLKD\VROHFyUăQJYӟLJkQJӕFOiNqPWKѭӡQJ
UӡL&k\QKӥKD\Fk\JӛQKӓ
&өPKRDNKiFJӕFFQJJӕFKRһFOѭӥQJWtQKWKjQK[LPÿѫQӣQiFKOi
ÿѫQÿӝFKD\WұSKӧSWKjQKE{QJÿѫQKD\FjQKFKXǤӣQiFKOitWNKLӣQJӑQ
+RDÿӵFFy-5 OiÿjL[ӃSYDQVӕQKӏEҵQJVӕOiÿjLQKXӷOpSKuQKFҫXKD\
TXҧOrKRDFiLFyOiÿjLGtQKWKjQKW~LSKtDÿӍQKOjPӝWPӓFy-UăQJQKӓ
EҫX FQJ GҥQJ YӟL ÿjL ÿҫX QKXӷ ÿѫQ PҧQK 4Xҧ EӃ QҵP WURQJ ÿjL Yj UѫL
FQJYӟLÿjLKҥWFyUӉQҵPWUrQ
Chi Boehmeria có WӟLORjLӣQKLӋWÿӟLYj%ҳFFұQQKLӋWÿӟLӢ QѭӟFWD
FyORjLPjORjLÿѭӧFELӃWF{QJGөng:
B. diffusa Wedd. [B. Clidemioides
Miq. Var. Diffusa (Wedd.) Hand. ±
Mazz.] ± WrQ7LӃQJ9LӋWOjGai lan.
&k\WKҧRVӕQJQKLӅXQăP hình dáng
Fk\OiPӑFVROHtWNKLPӑFÿӕLSKLӃQOi
KuQK WUӭQJ KD\ KuQK WUӭQJ KҽS dài 2,510cm, UӝQJ-5,5 cm ÿҫXQKӑQPpSFy
UăQJÿӅXKDLPһW FyO{QJQJҳQJkQJӕF
 FXӕQJ Oi GjL -7 FP &k\ FDR NKRҧQJ
cao 0,40 ± PFyO{QJQJҳQ

Hình 1.1. Boehmeria diffusa
Wedd. ± 1J͕QFk\PDQJKRD


-5-

+RDÿѫQWtQK FyWKӇ NKiFJӕFKD\FQJJӕFWұSKӧSWKjQKE{Qg mang
FiF[LPÿѫPWUzQ[LPÿѫP ÿӵF UӝQJ4PP[LPÿѫPFiLPP. Bao hoa 3-4
SKLӃQ QKӏ -4, qXDQK QKXӷ OpS KRD FiL GҥQJ ӕQJ GjL PP Fy - UăQJ
QKӑQYzLQKXӷGjLPP.
Cây UDKRDYjRNKRҧQJWKiQJ-WKѭӡQJPӑFӣGӑFVXӕLWӯYQJWKҩS
lên ÿӃQÿӝFDRP
ӢQѭӟFWD cây Gai Lan SKkQEӕ Wӯ/jR&DL6D3D
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YjRWӟL.KiQK+Rj1KD7UDQJ
.
3KkQEӕӣ1DP7UXQJ4XӕF/jRYj9LӋW1DP
7RjQFk\ÿѭӧFGQJӣ9kQ1DP7UXQJ4XӕF
OjPWKXӕFWKDQKQKLӋW
JLҧLÿӝF
B. macrophylla Hornem. (B.platyphylla D. Don) ± WrQ7LӃQJYLӋW- Gai
QѭӟF
+uQKGҥQJFk\Fk\QKӥFyWKӇFDRNKRҧQJ 3-4m. Nhánh cây PjXQkXÿӓ
to 4-PP/iPӑFÿӕLKuQKWUӭQJUӝQJKD\WUiL[RDQFyPNJLQKӑQJҫQQKѭ
FөWӣJӕFÿӝWQJӝWWKjQKÿX{LGjL-3cmGjLWҩWFҧ-10cmUӝQJ-FPUҧL
rác O{QJViWӣFҧKDLPһWUăQJKuQKWDPJLiFJkQJӕFPjJkQErQQJҳQ
KѫQJkQbên 2 ÿ{LFXӕQJOiPҧQKGjL-11cm.

Hình 1.2. Boehmeria macrophylla Hornem. ± 1J͕QFk\PDQJKRD


-6-

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