Các phức hệ sinh thái – tuổi diatomeae trong trầm tích holoxen ở đồng bằng thanh hóa – vinh
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- Cac pb~c b~ sinb tbai-tu~i Diatomeae tron~ tr~m
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to
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t~cb Roloxen ~ d8n~ bang Thanb HOB - Vinh • ••
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NGUYEN KINE
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ki~m b Vi~t Nam ••••••
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IJj; sum THh-TUoI
DIATOMEAE TRONG
TR~M TicH HOLOXEN 5 OONGBANG THANH 80"- VlNHzyxwvutsrqponmlkjihgf
'!'rAn~c Thtnb.D~ng ~o Ng.
(Del ~c TAng bgp H8 NOt)
D~n~ h-An Quan, Ngu.y~l'l
ti~n Cbu
(Doen 204-Llen ~oen ben ~~)
cu.ng nhl1 ~~ng bAng B3c BO, cic trAm t!ob Holoxen CI ~Ang
baDg
~b.nh Ro.-Vlnb
nhlAu 10,1 nguAn g60 kbee nb,u, obuy4n b~ng nbanb
theo chlAu tbAn~ ~n~ YB nAm ngong vl vOy, vite pb~n ohis v; 113nt~
obUng qua ceo 1~ khoan t~dng 461'kho kh~n VB cbI 0& tbA tIn c" ~
..
,.'vi 00 elat. Que pb!n t {cb Dlatonb~.phAn
t10b cac pbl10 bl sinh thoi
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mese V8 ngh1en eeu C80 tram t10b Holoxe.a ~'..
co
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O~c trAm tioh Roloxen 1~ ro tren dl,n rOn~ nh4t CI d~n~
blng
Thanh ~a- Vinh. ChUng' n1m khOng eblnb en trin cac trAm dch co tu!i
Plelxtoxen ho~c n3m trvo tlgp tr3n da g~o v~l b~ day r!t tb8Y d&l ~
I!t~tv~i~m8t d~n 4311 ( LK 14 ) trong kbl bA d8y l~n nb4t oua trAm tb
Dt tti IJ vUng nay chi 129fSm ( LX 13). Qua pMn dob chung tal
tbd)r3ng cDC boa tbooh tao Ddatomeae ~'p ~
pbong pbu trong trAm t{eb
Roloxen, d~o bitt Ie tronr, cac t~Am t{cb nguAn g6c,b14n. C80 phtto h'
sinh thai Distomeae ~Jp trong cae l~ trAm t!eh tbey dAi tbeo chlAu
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tb8n~ d~~ ~ CDC 10 kboon va tron~ cee tram t ch cun~ tuoi kb8Ct~
O'vun~ nBblen c~ t~n toi hei bt th~~ pbtic bf sinh thni son~ son~
tlnb t~nq pbet tri~h oua CftC dlAu kien cA ~J8 11
nhau pbsn anh ~u~~
tbeo kh8ng gian va th~l gien. Hf th~n~ thti nb!t g~m 4 pbt1c bf
pb8n
b6 CI nh~ng n~i trung t~ ~An~ b~n~
nh;n~ n~l n1m Bst bi4n
blfn
nay ( rIa dong ~~llg b'n~ ). Cao phtic h~ sinh thei i:u6i CI ~§y thA bitn
ro rong nh4t ~ 1~ thoen 14. H~ tb6n~ tbtt bo! g~m 3 phtic h~ pbAn
b6
~ nh~ng ml~n gAn sat nut, xa bi~n bi~~ tol vo th4 bi'n ro nh!t CI 18
khonn 6. TUAi vs
t~~ng tiDR gi~e eae phtic hf sinh thei tbuOc
b81
b~ th6n~ ~I1VO tb~ bitn trong blnb 1.
vo
sv
Theo ceo phtio b~ sinh thai Diotomeoe tuAi D18tomeae.Cbu~
t~i
00
ohio cae trAm t!ch Holoxen CI Thanb R06-Vinh tbenh c80 trAm doh
1
tu~i Hol~yon s~ ( Qrv ) ~oloxen gi~a ( QrV2 ) vo Roloxen muOn(QIV~
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1- CttcVh~ s Inb thsi Dintomene tuol Iioloxen SO'DI.Quotr nh thanh
too
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cac trAm t{oh Holoxen b ~~n~ blng Thttnh Hoa-Vinb g~n 11~n v~l
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ot! pbat trien cua blen tien Fll1ndrien vs anh htlo'ngclJ the CUt! no to'
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~on~ ph~c b& n;,.cae dong blAn cbl~m kbosng 4~ «to trU'~
1e.
Grog.
sp., Thel,
trAm dOh
cho chunG
d's
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cu6i
kho;ng
ki~n
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b~ tr(\n 41
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hon
c ht;c m
bOo ). O'n
ho~c
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C05cinodiecue
kton biAn
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lineatus
Ebr.
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Cosc~nodiecu~ ap.
donlcus Cl.
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b~
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Goml-'bOne1118tlcumiratum.
Melosira lslandic8 Miill.
M. Itellna ( Ehr.) Klit~.
C;yclotelll1_comto (Ehr.) Klitz.
Cy. Ste111~er.
Cl. et Gru~.
NitZ8Ch1~ ap.
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bgh~~nbrihs1 b~. 1
tb~n~ ( tr1~u dang, bao 19t...v.v.). 08C d,ng Diatomese 190 ~ J •
----5------------1
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g~9 sinh
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chigm b~n 90% tron~ do chu ygu 18 cec·doi bi~u tbu~c gi6n~ Eunotls
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khoc nh\l BDc111sria paradoxa ( Gmel.); Synedrs sp.Nltzsohia ep.v.v.
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, B!~n van bOt' r
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C1~ NhU' vty, tren cd~ phdn t~cb~dn~
be day va 40 sAu phAn bo cae
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tram t~ch Holoxen sam VD cac philo hi si~h t~8i Di.tomeae d~c trU'ng
Lito c1'~!1
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oho ohunG co thethay ranp, bien tien Flandrien V80 dau Roloxen sam
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~o ti~n VDO dAn~ b&ng Thanh Hos.Vinh v~l m~c nlldo x4p xl -4Om
~ijn ou6i Holoxen Sdm d,t t~l m~c x4p x! -~Om Vdl t~o
trun~ blnh
•
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kho8n~
2m.m n~m ( neu
nbU' col anh
h~ng ou. bi~n d~ chuyen
4~n~
8. 6'080 l~ kho.n
ki~n tfokhon~ donp;'vai tro quan trong ).
.
tll~
bi~n
2- otic nb1ic h~ sinh thoi tuAi Holoxen gliis
t{ob Roloxen s&m
8 oat tb·ocb sn.tl
oo~ trAm t1ch Holoxen p;iiio od bon Is nhiiDP;trAm t{ch biAnphAn
ã Pb~c bĐ D!,tob6 trEln <11&n 10 Idn nh4t tr~n d~n~ b~np; vdi b~ doy they dAi t~ 3-511
86 ltiVng(hCfl~beo
( LK 14, LK 13•••v.v. ) nh~ng v~n~ ms thc,l
nay do khs X8 bc,) dgn
• 1111&n
otis sang. •
hO'n ch1;c met ( 0 LK 6, LK 29, 1,K 25 nhii~ vun~ me thdl kY My 18 ven
Ie trllnp;
bc1 ). 6' nht;np:vun~ kha XI} bC,:'tho nh pb~n,.trAm t{ch la set xam xs nh
,
~o
- Ie
kY
Ie.
bo~c set bOt xam den. Ph~c bO sinh 'thai Diatomeae ~~c trU'ng
cho
ohung ( b ) co s~ l~~np; trunp; blOh va hAu nhU' chi ~~p 000 deng F1snkton bi~n kb~i I( tren 98% R~ lu~n~ co thA ) nhU':
.
bl~m 5%. Cac d,ng
~
v., bau
nhll to~n
blgn nb~t 1••
Ccaoinodiao~~ 1lneatu9 Bhr.
Coscinodlsous ap;
~hltscus Coclatua Bail.
Thalassiothrix longiasima Cl. et Grun.
Th. Fr9nenfeldil Grun.
Thalassionema nitzsChioides Grun. v.v •••
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- ndi yen b~.. c~' be•. day
, tren
Cnc
d~di cb~c met thU'~na:10 set b~t meu xnm ~en eh1io chi~u di t!ch thVc
~ot b~ thun bun hoa vo Y"C cae th:in mem bien( DentDl1um ep:Plocenta
op •••v.v.). Trang ph6c b~'sinh thnl ( b') ~~c trun~ cho cae
tr~m
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t !ch nny ~Ip t~i 60% CD the Plankton bien nhU':
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h1~u 1~
kboSD
V(1t bA d8Y
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T~o18ss108iro exoentrlc8 ( Jlbr. ) Cl.
Nitzscbia pacifioa Oupp
Tbolessionemo nitz8chioides Gru~.
Tbol8ssioth~ix lo~issim8 01. et Grun.
Th •.ProueDteldl1 Grun.yxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
000 d:fns Dioto eo ngu&n ~6c nU'6c n~t
va 1'g DlO m~t phln Imt
t~ l~c 4~D too ro 1 n~ dVns o~ng trla t!cb set b~t chl4m
kh08~
~.
Dfi difn
( C ) d~c trU"ngch
phong pbu v~ theM
g~c blin ehi~m uu
Coscluod1scu
COSt aubtil1
Cos. divlsus
COSt Oculus-
Is.
008. rodiotu
Oos. lneustr
cos', Curvat
Oos. lineotu
Coecinodiscll
Csmpy10discil
Amphora er,
Am. aff. b;ys
Am. ostreari
»tnoti8 grscislis
Eunoth sp,
Eunotie falox var. grocl111ma
1f1tzsol0 ap.
Staph.nodisous sp.
T.V.
S,
ph! bi4n ~n~ rai V8 t~dn! 46i 4Au kb4p tr3n 4~n~ b3~o"8
080 trAm t!oh Rolax n gl;a V8 oao pb~ ~ sinh thoi Diatomeoe d~o
t~ng oho chPng ob~
to blAn tign trong giol dotn nay ~ 4&ng bIng
Tbanh Hoa- Vinb 4 t tc1i t6c' 0 r4t l~n.onmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
Ii U th1ie nhf.n Il\,fc
n~do blAn
30a
veo ou61 Roloxen
~ -4!u 'Ro1oxen gi; d Tbenh Roa-Vlob d Il~C
e
va mv 000 nb~t mo 4vt b1 n tdo os,. 4ft dtfgc 18 4-5m- tbl v fJ 1
khoen~ tb~i ion keo dol cue Holoxen gi;8 kb08~ 2-3 o~bln n~m (t~
6-7 n~bln nam d~n 4 ngbln nlm cuoh ng~,. nay ) tbl t60 ~ doo~ Cue
biAn pb'i tren 1cm mOt n
con 86 no,. bttOo chung ta ph.i l~u tAm
dgn vol tro dong gop cuo obuyAn d~ng tAn kign t,o.
Am. me:dca
sp.
Trloerstlum
N:ltzschia ru
Nltz. punge
Nitz. pscl:t1
Nitzsohia sp
Aohnl1nthes 1
Aohnantbes s
Dentico.le of
Ths1sssiothr
Th. f'rouenfe
Tbdessionem
Diplonels sm
Navicull1 of.
Surielll1 fss
Amphora
Toy nbi&n. Tbee oao pb~o bf sinh tbai Diatemeae ~
blnb
thonb trong\~181 doen nay d 4ang bAng ohu~ tal kbang tbiy co d4u
hl~u ~l ch~ng to blAn ROloxen gl;o d n~l nO,. 00 nh1,t 4~ 000
h~
ml';o blAn ~nh p&c B~ b8.y giC,. Dfo bl~t. Trong oao pbtic h~ nay hoOn
toan v~ng m~t ooe dvn~ Oosoinod180t8 nodullfer. Oyolotella striate.
~loer8tl~~8VUS
v6n r4t phon~ rbu V8 d~c tr~ng'cho cao blin ~
nblft 46>1, ofn nbl~t f!d1 nblt 40 gJp tron~ ceo trb t!ob
Roloxen
gl~8 b 4~ng blng oong Cvu Long.
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,
1
3- Cao
ph~c b~ sinh tbei
tuol
Ro1oxen
mupn.
BiAn tlAn Roloxen gl;s sou khi 40t m~o ove 401 4 - 5m 80ng
Roloxen muOn r9~ xu6ng t~ t~ va 4'n m\,fc+~ thl tom d~n~.
Trong
..
hoon conh 4y qua trlnb l~ng dpng trAm t!ob~Tang Ten biAn ven vung
khao nheu.
rla phis dong cua 4&n~ b3n~ hi~n nay
co
O·vun~ ven bi4n bl',n nay vao tbC,i p:1sn ctAu cuo Ro1oxen ",~n
oon tAn tel cha ct~bi~n. TrAm t!ch biAn ~~p ~
b ooc 18 khoan13,
14 co b~ dsy kbo !6n ( tdi 22m
- t~ dO sAu 2 f!&n ?m ~ L(14)
vel
thnnb pbAn tb~~~ 10 oet b~t mou xom den, d81 khi Ie cot b,t oh~o
cec d1 t!oh thVC vOt vo xSc oeo tbAn m&m blAn. PbUe h~ sinb
the
8
A
Tronp: cac dO
thuDc p:l~n
ah~ y~u
bel cbI
tium
rr.~'v sO
It .•
favus
••• v.v •
b1~n nbiDt ~~1.~
Tron~ sA nny phAn
khnc d~8 t~ l~c d~
Melosira
gr!
v
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Ih
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C
d~c tr~ng cho cac tram t1~ noy rat ~ioU va So l~ng.ca
the t
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'Phong phu,,,, ve thanh phan
giong
1001
60 dfng ) • Trong
g~c bi~n cbi~m \IU tbg v~ s~ l~ng C8 th~ ( 70% ); phA bl~n 181
Coecillodiscus marginstus Ebr.
Coso subtille Ebr.
Coe. divisus Grun.
IV 11I6 m~tzyxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
pbAn lcm
Cos. Oculus-irid1s Ehr.
~'OtChi&'
kh08~
Coso rodlatus Ehr.
Cos. lacustris Grun.
ces', Curvatulus Grun.
Coso linestus Ehr.
Coscinodisous sp.
Campylodiscu8 unduletus Grev.
Ampbora cf. crasss'Greg.
Am. oft. byallna Kiitz.
tr8n ~~n~ b~~On8
Am. ostreeria Breb.
:ho1 D1stomeoe
n nAyonmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
d ~&ng b&ng
Ampbora
SPa
nh5.n m~c n\ldo bi~n
Triceratlum favus Ehr.
30.
s-V1nh (j Illic
N~tzscbia rupastris Pr-lovr.
4-510- thl v fJ 1
N1tz. pungens Grun.
2-3 n~bln n~ (t~
Nitz. pacifica CuPP •.
1 t80 ~ don~ cuo
Nitzschia
sp.
ta ph;i l~u tAm
AcbnDnthes longipes A~.
o.
Achnanthes ap.
0III8S8
d\lQ'O
blnb
Dentlcula cf. murine
kb8ng tbly co diu
Thalassiotbrix lonF~ss1mo 01. et Grun.
nbift 40 co 0
hC1n
Th. frouenfeldi1 Grun.
DC pbllch'~ Day hoOn
Tbslassionemo n1t~scbioides Grun.
Cyolote118 striata.
D!ploneis smith!l ( Breb.) cleve
oho 080 biAD &m
NavlcuJ.lIcf. rouxl01des Shesh.
,~ dob
Roloxen
Suri811a fastusss Ebr •
(
)
.
•• v.v.
u~n.
Bong
Tronp;
blAn ven vung
4V1
4 - 5m
rtom dtin~.
ITeD
u.
Roloxen ~ttOn
d CDC l~ khoon 13,
n '7m fJ LK14) v(h
,
,
It
i 18 cat b'vt chilO
CUD
Tron~ CDC dOng kA tren, cac dOl biAu b~nlot chl~m kh08n~ 5 %
chu y~u thu~c ~l~n~ Amphora. HBu h3t cac don~ th~ng B~n~ ~ blAn ven
be' cbi rr.~'v B~ !t :'!ongs!Sn~ b biAn khc1i nhU' Coso Cwratulus; 'l'ricerlltium favue .••v.v. Trong
t~ng·e6 C8
tbl.
Trona: s~ nay phAn nbi~'l ls CDC dOnI':~~p ~ ml~n ctia sang, ~t
86
khoc ~~8 t~ l~c ~~8 ra I~n~ dQn~ cun~ trAm t!cb nbU's
Melosira grenulota ( Ehr.) Rolf.
I
hlic b, sinh
th "
9
M, 1 10ndiCD Miill.yxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
e
italic (Ehr.) Kiitz.
C ot 11e co ta ( Ehr.) Kiitz.
Cy. St lli~er8 Cl. t Grun.
&ntzscbla
p.
S n dro ap,
Syn dro fOB iculoto
notio p.
E folax vo~. gracillimo
E. 001011
Go pbon me 8cuminatum
F 0 11 1.0 p'.
Fttbellorio
ap.
Ep1th mia p.
Epith 1
bra ( Ehr.) Kiitz.
tb la turgidD ( Ebr.) Klltz.
Cymbells sp
Gomphocymbe
Tobellarisonm
f
Tabelleris
Frol':ilaris
Peronie sp,
Diatom sp. '
Svnedra sp,
Sy. tabuleta
Sy. 4cus.
Sy. ;iffinis
Nitzschia ac:
Nitz. cr, ro
Nitzschia 8
v.~ •••
bttc 6 Binb thoi C'n/lUoD- g60 l~o
~n. O'nhUtng nC1iv n biAn hien noy C90 trAm dob
tr n 0'0 tr
t!ch bi n Holoxen m~On ch~o ph~ hf C oon ~
l~ hoon
phis tAy 4~n~ b~ng ( h& tb6ng slnh the! tb~ II)
ren cao trAm t!cb chttophUe bf ~'( b1An ven b~ ) co nC11
trAn tAng vInh Phuo ( QIII' ) 00 nC11, oec
trAm
o U ~
y 10 thon ph n
uvl, 00 nb;ng nC1ith!y.ro bal pbln. Pbln
c t
t, 't bo 0 ost mou th~dng xnm
xnc c~v co rn~c n't con pbAn'trAn 1s cec tr~m
t!ch
1
t ma nAu gl,l,
ntluvang , xom v~np.:,xam tr~n~. M d8Y 1~
t c .•
cDo t m t!cb 11,\0
at
o h~ lob thoi C'
bent't 0% chu y~u thl\~e 17,iAng Eunotio sang ham canb "
b~
cbu y~U
Qn
IV~ :
g&m:
I,'
IVI
otia
Eulle;t;1El
'0
a
I>,
III ~ :
p:r cial is
•. altlx "or.
~rac
on roll sp,
h mia tur~l
( Ebr.)
Klitz.
YUtz
Thw.
b niD curvata ( Kiit?.) Grun.
ria ct. paradoxo ( Gmel.)
o 10 ~ibbD ( Ehr.) Omiill.
010
s1.Ii tbH
(Ehr.)
n. 2.
SO' ct~ bi.
bi ~n VD
(
A
)
va
n : % d
/
Cymbells sp
Gompbocymbelln sp.
Tnbellorin fenestrato (Lyn~b.) K~tz.
Tobellarill ap,
Fro p;ila rill sp;
Peronie sp.
Diatom sp,
Svnedro ap.
Sy. tabuleta ( Ag. ) Kii tz.
Sy. 4cus.
Sy. itf finis
Nitzsohin llctinostroides
Ni tz. of. roc ts
Hantz.
Nitzschia sp •
•. v. v ••
trong 08c trAm ticb
y cac trAm dob
pblie b'~ Coon
;,
sinh the! thti II)
n yen
net!,
bOt ) 00
1'1~1
-:-.-::-:
___
i.
rL~
!-IIJY
I
II
- - .
2t
.=' -. -:...- ~l
I~:!>,:,:
b~,
~
C>
- --
.,
' ••
hi
IV<-
,[I
I
I
:.
#.
- ..
-
-
+tr
i
"
1~.2
~ 1-4
,
- .
I d;~
ifiv'
=.
'-
":--"';-:.'
..
21.'
19':
II~
.~oJ~~
~_
t
fi!TIf±fJ
.
a
A
bi~u
di~n tt1(..l/!:
.
b;~n v~ cnc rlonp: nr;u~n
8~
~~&
so "
B
SO' ct~
,
--
1113 i
B.2.
'~ mll1ii
Ii'idi,
.. -.-
t'".d++++-+++H
-
b
..
J
It
!
I
" *;1:;0-.-,'
...
-.
.'-'_ _ _
phonp':
ben-
c,
II~::
~,I~:~~
I iam~~
'I'
tr3nlt. M dey 1001
tllC1ng
t!6i
, oao dO~
~ belli canb
I,
QI
.- --.
=i=
C
.
.
1V3 .
2~
C80
( LK 6
ty ,~ 7.zyxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPO
J•••n ;'"e :!
h"onmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
-- •.. '10
In.le
~:;It~~~~
~. I ff/lf.rl t,1
::.::' i ,I',
na!
trhm
bsi phAn.
PbAn
bttOc t1t<1ng b~,
deb
• cae trAm
o
t;l~%
rt-khijc.?tJ~~~
t:r::I]
b
n ty IQ ",. ~ii1a~cac rlon~nl7'u~n
~
p;6c nt10tc nr:~t - JQ' d l~ khoaD
qua
R'OC
14-
( Ii ) vii 15 khon n 6 ( B ).
p
:
%
,
CDC
J
dOn~ bien
"
~ b : % cac dCnR'
D~dO
nR'~t - IQ'.
11
Tom 10i b ~~n~ bln~ !bonb hOD - Vinh, trong CDC tr3m t!ch Holoxen t~n toi h~i h~ tb~ng sinh thoi - tu61 Diotomene. QU8 trlnh tiAn
hoa cua hai ~f th~ng sinh the! ney co lien quan cb~t cha tdl ' 8 n b
hllc,np;
CUD bi~n tl~n Flondrten b3ng nDy. N~hten cu-u chi ti3t
cac ph~ b~ nay cbo phep xac ~uen g~c tr~m t!ch,
sv
vo
l,
cbuyen
t~ng
"
CUD
1
chung • Trin C~ s~'.' 6y co theI li~n kAt VD, chung.
tlAn boe cua cec pb~c h& sinb th81 CUD bai bf tb~n~
vU8
n~u thA bitn cbu y6u ~ t~ng quon ty 1~ gl~8 oao dOng co ~u8n
ReC
bi~n
080 d,nR n~uen g5c n~c lV, n~~t ~
trlnh bey tren blnh 4
Sv
ve
~c
An
Ou3i c~ng , Oeo too ~!8xln ohAn thonb com ~n gop y aho ~ol boo.
dO
vo
LA
A. TRAM IOUO
"
vINzyxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
Lloo
OHINHonmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
•
1. Allen W.E. snd Oupp B.E. 1935
Plankton diotoms ot tbe Java Sea.
2.
3..
4.
5.
6.
7.
12
Tbe Sorips • lnst of ooeon uniY. O,lifornia
Bno 080 d~8 ch3t t~ B8i Pho~-Nam ~nb
Thanh Ho.-Vinh
ty If 1t200.'000 1978-1981 lllu trti T6np: o~o ~8 oMt.
Crosby l·.H. ~lnd Fevgtlson Wood B. J•••v,v. 1958-1961
Studies on Austrellan ond New Zealand 4ietaaa vol. 8588. Bull. "new zellltlndoc.sn. inst. 0011. nap.
Cupp. E.E. 1943
Marin Plankton diatoms of tbe West Cosst. ot North
emerics. Bull. Sorip. inse. Oosen. ot. univ. Csl1for nie.
Vol 5. ~ 1 •
Hendey N.J. 1964. An lntroductor,y'occount ot the Smaller
al~8e of Britisb coastal Wster. Par. V. Booi1ler,vopbyeae
(Distoms). Fisbery Invertogstion Seve IV.
Hust~dt F. 1927-1930
Die Deuschlsnde, OstevrOlobs uDd dev Scbweis in , R~benhovsts Kryto~~mer PIors IV. T.1: Lelpreg
Juze A.P. 196'. Nghi~n e~u ~~s tftnp:V8 cA ~~6 ly. vUn~ tay
b~o
Blnb ~ng
(theo Diotomeoe).
NXB Vlfn &11 LAm KH. L!8n %6
(TUng Np:s)
va
~~ol
x
'cBezyxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
trb deh Holo1'1
e. Qun tr oh tl!n
·
8. Klm We, TtI<1ng••
v,v, 1967. Trunp: qu5c b~1.,d\l(1ngPba
du
yxwvutsrqponmlkjihgfedcbaZYXWVUTSRQPONMLKJIHGFEDCBA
che tot!
1I n h
Kbuet~o 100i. Tb~np: H8i xuBt ben xi. (Ti3nq Truh~Qu~c)
~hien c~u chi t13t
9. Khurxevich G.K. 1976. L~cb s~ pbet triAn b~ thvc v,t Dis~Bm t!Ch, va
.8V
tomeee ~ b8n ~~e b~ Nerocben. Minekd
10. Kokubo s. 1955. Pbo. du Khu3 tso 10fl (TUng Trunp: Qu~o •
kAt VD ~Jnh tuAi
~i 1960)
D~oh t~.t18n~ Nb$t. Tb~~
W8
ai bf tMng
11.
Mubine
W.
1974
fllg
~u8n
p:~c
h bs;r tren hlnh 2.
Problem of dIatoms and Sl11cof1ap:ellatee Queternary
- Stratigrapb;r In the Equ8torl,1 Pocifl0 Ooe.n
.
: <1nI!~ng cb!
L&
12. PbAn t!oh Dietomeee, 1949. T.l. NIB. vln 11ta ~~e oh~t
qu~c gie MS8odVa.
.,t
I
_..
•
co
ornh
i Thanh
cy'c
Ho.-Vinb
cb8t.
~8
13. Prosklna-Lavrenko
A.I. 1963. Teo Distome,e bem ~8:r ~ BiAn
Den
- Moo 1.
14. Teo ,D1etomeee bo. tb,oh c, LUn xe, 1968. ·"Nellk8"-M.
15. Teo Di.tomeee ~ Li3n X8.' T.l. 1974. M-L.
16. Sbirota , 1967
The Plankton or the South VietNam. JspaD
17. Schi1 tt P. 1900
(Dletomeee) In: En~18T A. and ~- Prnntl
( . Beci11arlales
"Die natl1rl1cben Pf1snzenfsmllien ••• " Pel1X;AbtI.Lelpri~
18. Xksbiobavskli A.P. Teo Di8tome~e trai nAi n~e n~pt ~ Lien
Xo. Moskva. Univ.
1958-1961
dletoaa vol.
11. Dep.
85Ng~~ nh~n be! 8 theng ?nam 1982
at. of North
aniv. Celifor ••
t of the Smeller
• Bocille~ophycae
I~
cbweis in , Rabene~
&
o
,.
dla 1y vun/!':
M;r
Np:, )
13
