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NGHIÊN CỨU CÁC ĐIỀU KIỆN NUÔI CẤY VI TẢO BIỂN DỊ DƯỠNG SCHIZOCHYTRIUM XT41 ỨNG DỤNG TRONG THỨC ĂN CHĂN NUÔI GIA CẦM

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Khóa luận tốt nghiệp 2014 Khoa Sinh Học


  !!"Phân lập,
sàng lọc vi tảo biển dị dưỡng họ Thraustochytrid ở rừng ngập mặn Xuân Thủy,
Nam Định” "#$%&'(()*$%+,%
-./0123)45'67894:)7:
;/<)=>' **?3<2678@=
&3%ABC?:D) 6E;/%=
&FG<' H#A
3)4'I,J5', %!;)<B67
;/<)=
&' H&*K2 )%$#%
&'()$#%*+3'I,!,L;E9M
,!;)<B94%)7:!=
&' N/4O*?P K,Q 3<
Q67894:;/<)2.=*?
,R S<,!;T H/
E=
012UV WXYZ
$
+4&A
,,
Khóa luận tốt nghiệp 2014 Khoa Sinh Học

2.1.1. Đối tượng nghiên cứu 12

!"#$%&'(##&)*
2.3.4.2. Phương pháp tách acid béo 17
+*
!,#$%&'(-.#(/#&)0


3.4.1. Xác định lipid nội bào 28
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
,/0123
*L &[
\]^ _ R/]^
\]` _ R/a`
OO O/I/I
b0O b/R/R/I
b"O b7R/R/I
P"O P/7R/R/I
*"cdO
*/"<//RIe//I]OIf;
/. ,I
gcdO
g//RIe//I]OIf; 2.

0"*
07Re /R<9I //7 ] $;M <
/7
"*K "<<7R<
"cdO
"<//RIe//I]OIf;/
(&Kbb ('SI?IA8
h1g h@)7 i
&j &%<AB;
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
,4
 !"#
$%&'() *+,(-./01234567"
89:;<=$%&'() !#"

5->(?@A&()&(B/CD,(()1=$%&'() E
7$FGDH,;I@A&(JKL7?@A&()M(AE
7$FDNOPQ?D=(?@A&KR7(A,(S;I@A&(JKE
T7(A(?@A&-> DE
8U$;V*,()OP WX
8G/YO>2A(?@A&=9LRR$%&'() Z534567[
8\M=D;?+()M=$%&'() Z5
88\M=](;I;?+F+()M=$%&'() Z5
85\M=V;I ^;?+F+()M=$%&'() Z58
8X\M=V;IO&+W;?+F+()M=$%&'() Z5
X
8E_<&+()M=9LRR$%&'() Z5()&](^@
OP W6O0("
8[?@A&@`( A2A3(),72A(?@A&()./I OW)W-3DG7=,
>(OD-()&(?@A&9LRR$%&'() Z5E
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
,5
LGAD ()<a'
LG8A ObOD-(c+^2A()dOb/=9LRR$%&'() Z5
DNOeD(BfgZN='K _h6
LG8\M=D;?+F+()M=$%&'() Z56
LG88\M=](;I;?+F+()M=$%&'() Z5
LG85\M=V;I ^;?+F+()M=$%&'() Z58
LG8X\M=V;IO&+W;?+F+()M=9LRR
$%&'() Z5X
LG8UW&-i*,()OP WU
LG8"AD-@j&=9LRR$%&'() Z5[
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
678
1/;327S/k!!k<AB/

:7D   2?l9/%=&)/<m:
'k;I/V/ #Q,!k!<,=
&EVVABGInk;llVAC RV
%6DACS,/L'7D o,o=&R 2
<AB<H/k;QIA'7D V,
 %E;RAIp=(4)[24 QI
IA8/qEVVS' 'k<AB:7D <
p=
&R! E '?GIn;H,:
7D k;EVV=('.2AC/Vk
,;./IrQl,/Hk7i
kIIA8;'V///7</If;/.
  s"<//RI  e/  /I  ]  "cdOt    /I/R  C  I,    2
<R7<//IR=&.;'ABGIn< EVV
<'SI?IA8s(&KbbtABGIn23=g2.<(&Kbb'
k  ! \]^  Au Thrautochytrium aureum Schizochytrium 77=
Crypthecodinium cohniiUlkenia77=Amphidinium carteraeLabyrinthula 77=
<Schizochytrium'<ABb0O/k,Y^v w<w=gi
;GIn(&KbbS'kb0Ok!AS AuIrI
I4!;k.A;?'ACC p/n;l
)S;S AB94'kk<AB'7D =(4)
<%AB''kEVV<E9/%= &
?IIA8<,'/qpR!; A
.,'A22 i=>k7@:6
3E!u"+9:;<=>?@&ABC
DD(EschizochytriumFGH9DI&'&9JJ?#K".
1LI;!?!;.ASchyzochytrium AC
7S=&ABB7k9/%qEVV
A"cdOO//=
+4&A Y

Khóa luận tốt nghiệp 2014 Khoa Sinh Học
(MNO5P8.QR8
HH9JJ?#K
HHHSS&9JJ?#K&1>&#
&YXV VAH/7S ,H.2V
AC`xwV =&R.;[/"#&EVVs*n*V]K2
117"S1t/,WXXW]WXYY EVAC
4[/'<ABEVV7,;'
Y^UxwV =+< 2 E @:7S[/V
V7[/AH/yYvz=

04Y=Y=*V/ yW{z
1<lS'kEVV<,;/
 @{X]vXx;.<ABEV|B7 EV7R;
k/ / /I WX]^Xx;.<ABEV|B7Q <,
2A=&:Qk7<EVV
k776/Q '7D ;/Q'7D 7n[/
7['=&!.'AC;'V
E[/Q'kEVVAH=*nS
Il; C2 )AVkk7;
9}k/Iln; C2I</2
V/=*l4)!V 9/'kEVVAH
/7'7n2!)7;D=&R*n*VV WXYYAH/
)7;Dk75v=~k<EVVV^<HV
+4&A W
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
WXX`=& EVV<ABA<6/ •  •<^=v`
ks ;'Z^xtEV, A|A;I<,2
2?A=== Uks;'UZxtEVq;'
X=Wks ^xt=hvV WXYW'AH3)7Y=Yk

Y=vk<6/ •=(7n29!<)7;D
;'k?D<9/I/727'!
<!5'7D ;?AEV<n2
yY~z=
g2L'779/%<4 /Q< Hq
p<6/'94,7n2Q<)7;D=
([2.A/Q<AHS'kk7
QEV7R<=
bGIn7REV/  2V/
!QIIA8@(&KbbH <ABI/R/R/Isb0Ot
/Ifk<AB//ABR fSq2V
/ =
HHTUDI&ABCDD(E&'&9JJ?#K
*k!<9/E;R/:7D /7<:
4 ; <,<. C/ 2?A2<H
;.'=1'IIA8;q'EV#
S /<,E;R2)=+<<IkS,
9/V I6Sk7Q<?AEV
Vq?/=
('<)[S'k2I7R/ 
.kL=*67S/!HQ4
;GInV<ABr<=&;ABk7!
QV<ABA |;.[/6€SVk7^i
Z<=&'S7SRk'AH.lIlQY
R/Q'S'kp 2<AB7R 2V H
WYR/)iZ~R/yWZz=
•!;7S. C':IA87'7n
+4&A ^
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
2i‚QV<ABS.?*_

W
,Qk7
/=&.i77' 2.ABS <HAu
;;!;7.kIrr <,)
;/)7pI=b<,(&KbbH;'VS/
/qV<AB@B7kL'7SH.<AB
;/94< R3ABR fS'k;.<HG
In94< RS< ' 7lQ=ƒHl7l
'kY;;.(&Kbbk7UI<</H7l'kC.
9/%.H':IA8yW`z=
(&KbbABGIn'kb0OA ,ACryptothecodinium
cohnii2Dinophyta Schizochytrium 77.,2Heterokonta=
(GIn<;/2 Thraustochytrium  Ulkenia 
HeterokontaHp 2 nl/ABE=
&IQ.(&Kbb<<.2;.[/
;k<AL=1<IA/7A777<)7.A
L.[!;7#l 94
A/lB7S' '/;.@./=0
/C(1/ [.'GInEVV
[Q.)7,94QA/l
B7H!;;l)AH9'R <,A//=b)
7<)7?<,GIn'< EV€i7!;;VA/;/
/#[/'FAA/7EAB:=
HHVKWUDI1 Schizochytrium&'&9JJ?#K
$GIn'< EVV2)AB@L
V {XAHl;'^Xx'<AB'HABIp'
kEVV=g2.E35/kf(&Kbb
/Q u<R<;'VRR//If; 2.;
 9/%iA\]^\]`=1/'<Qk7
/ AuOKYKWK^KYW*P/Ie</I

//I.<7<s X=UxX=Yx%<AB
;t/RIs X=YxX=Wx%<AB;t=&;D7V
+4&A Z
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
[// SGInU]YXx;.'/ 277R
A=*kf#k7/IfA/I<<R/I
<<R/I//I=*kf67#/// ObPj
.SSIrk7< I/ 8V [/<#
E=N;AB„;.'4iS A [/
I/FA<#EAB';SyZz=
1/kfEVFIn< ' 2n67'
!Ak7=jqkfEV6Mq
k./S'/f;'/ 
EV=
('SI?IA8Schizochytrium < 2<:/%/<MAC
Q\]^!.sIt2V?=&R!
Schizochytrium;Lb0O #;E/k2,F
A;;'V=*7l/k[
Schizochytrium;:k[/<,2.'AyY{z=
0 <AB"cdO]^b0OK<RV@W=vZ=X<;
H2VqW=WxSchizochytrium=+.H …E
  2  V    q   Schizochytrium H    <AB  A  A  Y`U 
b0Oww <ABb0OEV<Y^U wEVk7U<
HEAsWv b0OwEt=1/ <ABb0OE
SV<WWX wE.HVSchizochytrium AAH
vWU b0OwwyYZz=
HT1&ABCDD(ESchizochytrium
HTH1&'XYZ
&/I< 2 )S;;'V
9/  B7  /  Q  '  u Althornia, Diplophrys, Elina, Japonochytrium,

Thraustochytrium, Schizochytrium, Ulkenia;'^U<y`z=*6GIn
:VACGIn/Sl<F/If=
&AH;/&/IAB7%k C4;'V.I?
IA8F4A..H%k V
C<,I:/;9'E!RR7RA/3.
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k„&/IAB7<,';'VI?IA8
iAC:I[/9/R/RRR R ,<AHR7</ 
H<H7 R<<<R=(#[/S/Q I,
AuIA862G2GyYXz=(?
&/I[I:/4%C/,&/<<:;/
947'7G=
N&/I3AB6MI;'V'k 2<AB
<H/If\]^/Q uI/R/R/Isb0OtR/7R/R
/IsP"Ot'//I!9/%.HE;RAFA
Q['=&q<ABI[/&/I @YX]UXx
;..b0O ^X]{Xx=&/IF'k
~Xx/If; 2.sgc//RId/OI]gcdOtA
7/< <R/Is*Y`uYt<R/Is*YvuYtRR/Is*WXuYtR/I
s*WWuYt=
('SSchizochytrium< 2[/%&/I=&.
7<,%Schizochytrium AB77A/yYUzu
5L! * /s$/ R7</t
! 0RR;7/
LX! /<R/
[! &//<R
%! &//R/R
! Schizochytrium
HTT\;CW%]#&ABCDD(ESchizochytrium

SchizochytriumAB4 k!H@pG//H
S<<,IA8sk7nkL#//tAB7<)7[
 'nLA< npS)7 i=$;.
[/6S<H;'W=U†YX
v 
w%<AB;s&jt
;<9/ ';'V2.=
Schizochytrium I,4#)7@n  ,<AH
,kk7@! i;lAHWUY{‡ =Schizochytrium
iAC:7<n[/@R/I/I'
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k2G/ 7H4I,kAyUz=

04Y=W=Schizochytrium9/IAH;lSs†ZXt
-47S[/'AB„‚'7
2G=&.v/;k2G/ k
QV;lAH=*pH94V;lAH<:7/
[/,/L H=*7n
AC7S2Gi IA8y{z=04
Y=^ '94'[/%Schizochytrium
04Y=^=(#[/Schizochytrium yWYz
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(#[/ Schizochytrium/Q ^;•7SyWYzu
*;•Yu/,AC;,AB 2;l
AHk?6S42G'72
G=gi;6S4I,/ 7i7n
AC/,ACQ:947/7R=
*;•Wu7n7/v/AB'

7,7nAC,AC
*;•^u2k[/ACAB
,2G=$/ 2/2GAB'7
7SR;4;f7;l=
04Y=Z=04I,@;•7S[/Schizochytrium
Ot$:'72GKt&AC
*t$:7/<7[/bt0462G
Pt04I,/ 7
HTV\;C&ZXD
1E!4%3E „&/IE/,
<7IIrI9/AB iE/ <ABb0O/=*,
+4&A v
A)
B)
C)
D)
E)
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
<7I[/Schizochytrium <)7B77R<7I/I,!;lAH7
Q.7,=*,/#9/%
<AL)Skf'2:)=giIp,
<7I 2.<&/IIrI)kABA6/
kl!:4il[/6y{z=
AB<7Il<F)SAB/„k7
IA@//;,=
HTGXKDD(E]#1 Schizochytrium
(&KbbSchizochytrium <AB<7I {Xx%<ABk;
b0O ^Uxq./IfyYz=&A:[/R/ˆyYz
b0OsWZ]^Uxq/Ift/I7/< s^~]ZWxq/Ift<//I
f  !  k      [ Thraustochytrium 7  <)7  @  '  "//

"<77R=Schizochytrium7=sO&**WXvvvtF'k 2<AB<Hb0O
s^W=~xq/Ift7/< /IsWU=Yxq/Ift=&E
;[/R/ˆb0O ^Y=U^x7/< /I ^Z=~xq.
/If[/[S. Mangrovei‰O_]Y=
Schizochytrium   ;'  V  q  B7  2  <AB  <H  "cdO  Au
R/7RR/  /I  s*WXuU]^  P"Ot  //I  /I  s*WXuZ]`  OOt
I/R/R/Is*WWu`]^b0OtI7R/R/Is*WWuU]`b"Ot=
&P"Ob0O<//If\]^9/%y^z=P"O<B
,3ABGIn!?L/2 , 7<
A=b0O< 2/IfAFA2)A
.;</Ifl;[/3Š ,
=(4)b0O/#9/%:7S[/3Š ,
:4//,I4EV3.9
4AC=1/b0OF'ACl:!?
!  ,Au  ,29JV7S.
H'.<,;A klO<‹R RŒ ' =
*R/IA7/</I7/<<R;RRQ.@
"cdO]^Fk9/%:7S[/…….
<,:! , L/<A="cdO<Lkk9/
+4&A ~
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
%<m:IAB7D kIIA8y~z=
0/I<Qk7"cdOl=&Qk7
<H,=1/k!!/I3AB!kC4
S? r ;SC  k2 A; <,i7<</RI
7R<I]<LkiS H7nL //…
R =•In[/IA 2k7n/:7D ?,Ik!
<9/HiS  p?;?lq?/;f yvz=b
EQk7"cdO/"cdO@I3/AB
i9/ =

*(&KbbSE/ 2<AB<Hb0O6ABR <Q'
kb0O! V=SchizochytriumABR <<;'V'k
b0O<HC <AB<7I {Xx%<AB;q.
/Ifub0O ^Ux7/< /I WZxb"O Y^=`x
/I YX=Yx=$;.'FilB79
4,/If;/. ,Is*"cdOtI
67kq?=*"cdO@'k;<RR<
;?r ; <,i/7<<7R<s"*Kt=== pIr?
yYz=
1/ 2./RIk? i<///FAB4
kSchizochytrium=
O//< 2<,/RI 2I,.#//kf
/ /E/%<^^Œ]II]ŽŽ]/RR]ZZ•IR
 i:)<)9/B7;A.'k /
;D=1ABq:7D GInA.2 
/  yYZz=
Schizochytrium;'Vl<F`=Y w<///Zk
C AE/YXx<RQlX=^x# A
E/Wx<Rschizochytrium ;'V'/v w<Ž]/RI=
g2 'kAB$C1)K'3GInAH'[/ 
SSchizochytrium )k6S,/{={ w<///yY^z=
+4&A YX
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
HT^[&W_+9<&ABCDD(ESchizochytrium
•(1/ V WXXU(*%37<)7<,
'SI?IA8Labyrinthula7=Schizochytrium7=, 2.pSC
(  1/       <AB  b0O  sb/R/R  *WWu`  ]^t    b"O
sIR/7R/R*WWuU]^t/k7U]YX<H<,'S;=
g2  9  4        Q Schizochytrum I      g/R;
KRRsg}t<)7HkWX;.';w<l/Zvk

 <AB"cdO]^ YXx;.=
•1)K'k< R[/[Schizochytrium7=$hY,ZvY
%<AB;w<lY^^b0Ow<l/Zk=
+4&A YY
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
(MN1`R81Qa5/5bc8
TH1d&Z>+9
THH_&(e+9
*['SI?IA8Schizochytrium>&ZYAB)@"#$%
&'(()*$%+,%-./012
THT:fDII
$GIn  Inn,7#$%&'7#l
 2(()*$%]+,%-.N/012=
THVf#@&
*/kABGInkE@gR;s+Et$ /sg}t&
-.
TT?&'()?@
&E<:/ AkN"•q W
<,;O 7<<$R7 =1AHSIpS7/ A<k
@h1g>&[]1/ +?k<ABAH;/q/
l =gAkABpCYYX‘*WX76
iYYU‘*YU76=
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K'W=Y=&7 Ak
&7 A 0 <A%sw<t
N<R W
"R7R Y
*/k  R X=U
1AHS Y
TV(MX:X+9

TVH:X(MX:Xg:;W*W&'(h]#&A
TVHH  (M  X:X  g:  ;  W_  Z(e  &i  B  &i  B  Bj  Bk  ;i
lB#l'
"A77 „Q 1R/RyWWz=*k,Q
1R/Ru
KQ 1R/R< 2k [I;'^ AB/< W
+4&A YW
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
7=*7VH7L/CW3I%="L/AB/
I 23/k7/7X=Y ,//V ./=
g|V 4AB/Y`<H |<H<,AB/
Y` | 2Il<YwZXX
W
!/<YwYX =
$.<ABABlREu
b’s/w`Zt†YX
`
&u
bu<.<ABs.w <t
/u<.4 2Q
Cách tiến hành:
KQ </ RAB</,„Qk ;AH;
€'=/ RABiQ /;4k
://C?l76L/Q </ R=$/Ip
77RR"/R6 2lI?E/ €3AB<!AHk ,[/
</ RI?'‚<Q =
KQ E/SchizochytriumABA/<;lS9/C
);lZX?;lYX=+.HL € 9i;SlAB
4/R 7/<3AH; .7/<3;l;9'=
TVHTF:;&'%Z(e=?

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<%;;.'3 )2=
*u
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Y“ t3AB?%<ABAB;.H 2  ;=
$/k;2<%CYXX‘*Z=$/Zk;.^<<
7;%<AB;q=
TVT+9:i&_A(h;iW&'(h]#1 Schizochytrium
$:7S[/[Schizochytrium AB?R..u
2702kQ2<RQ2 .=g|.AB
;S /C E2;/=
TVTH(h]#X
+4&A Y^
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
(&KbbSchizochytriumAB<4/ WUX <H.2
<WXX#w76 AN"•C2Wv‘*Q2 .Y{”
<R<Wx=$/;k.'..<ABYX
U
w <ABG
Inl S?;'70lB7;.
'=&l AB:HI'70uZU`{v~=$/^`k<k
€?.<AB&jR7A772.3.1.1 2.3.1.2
TVTT(h]#>&;[
$/;?AB!;70lB7?2l
B7:7S[/Schizochytrium.
12'AC:AC7S[/(&Kbb=+S
? E2'AC[/2H;'VAC,;.[/
Schizochytrium<4/ WUX <H.<AB
.<YX
U

w <.2WXX#w76 AN"•C70lB73
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WUWv^X^W^{‘*=$/^`k<k €S?.<AB
&jR7A772.3.1.1 2.3.1.2
TVTV(h]#k;[_
g|<,'2? i;/u<;'Vl
H2 i/<;'Vl2 ik7=+S?;'2
i.A;.'l AB:HI'2 iuX
UYXYUWXWU”=gAN"•H7023AB?Cmục
2.3.2.1và 2.3.2.2<4/ WUX <.2WXX#w76
<R<Wx=$/^`k<k €S?.<AB&j
R7A772.3.1.1 2.3.1.2
TVTG(h]#k;[ZWl
0 <AB<R'AC;'VAC;lAH[/
(&KbbSchizochytrium=+S?Q2<RlB7:7
S;.[/Schizochytriuml AB<
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2.3.2.1, 2.3.2 2,.2.3.2.3HI'Q2<RuY^`~YWx=$/^`
k<k €S?.<AB&jR7A772.3.1.1 2.3.1.2
&@l ?'AC[/702Q2 .Q2
<R?AB!;klB7GInS<
R;.7nnE7R=
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Khóa luận tốt nghiệp 2014 Khoa Sinh Học
TVV(MX:XZ+l&W=_&#&'>&_BSZ+lHmZ&
TVVHnBo:&'SZ+l
P&'SZ+l
04^=`=$Q94< R
• nBBSZ+l
K4< RIlYX<l„[27)u;k

:70:<ABR#//:2An
;lA;l/<k €=
• nB&i&B
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?9/AHuG/,k;/pCWXX‘*`X76<77
;Gp„/</RCYWY‘*`X76.p<7i5
.=
+:70R#//2ABD?9/AHu5
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;lp

Khóa luận tốt nghiệp 2014 Khoa Sinh Học
< R=
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YX
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TVVT+l
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YX=X^†YX
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TVGX(MX:Xg:;ZXD&:#DBp
TVGH(MX:X[Zl'lD

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4Al<F< R[/77<7I=g
;9/:7IAH;lS•9/7n2:A/AH
[/ A9/=(7f72 1<RRI2 '/I,
kfls t77<7Is / t=
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I? 1/]<i„ Ak=-/:/q•
9/,AHZUX GIn;lS•9/O—R‰ /R
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ZX AH.[7is8Y^ †YXX t=O<;/</„
Y=X <k7'E‰sk7'E‰Q YUI IIRs1/_0tUX <
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;sPR</IR /gN]WYXXtU76=$/< !7p
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7AB< <,IAH#AH ,27#=gR</II?
O<;/<HX=W <k7'E‰‰s`U <`1I</IUU < R/<
0"*/IRt=$/; R</LCvX‘*Q[;'YX76
AB< <,IAH#AH C27#=bI? R</AB
qY=WU <[/kG‰‰‰sQ UX <]R/R] R<R<RR
YuYwtAB2!„ <W=XXX#YX76=b?H7l/
 AB  <, =  bI? AB  q    ^ <  .  G  ‰(  sY=W  I
IIRYXX <g<<]-tAB2!„/U76= C

W=UXX#YX76=b?pABS/.[ H
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2I? €AB< /HUX‡<IR<RR^XXsi/R]^XX]
R/Rt=&2 €ABk &*QABiQ[E/
YXX <]R/R]IR<RRsZuYwt^X]ZU76=OIf:;
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IR<RR^XX=&2!II?</R< <W=XXX#YX
76< W=UXX#U76iYX76=bI?AB/„
R/A;=g€AB#//C<,HUXk7'E
‰‰‰=bp77R<%N*6)=$/R 7l 
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Khóa luận tốt nghiệp 2014 Khoa Sinh Học
(MN$2P81Qq8`
VH. P#W:&SD&iBA] Schizochytrium FGH
*[/;ABk AN"•^`=$/
9/4;D<,m/,4I,IAH;lS
C27,s†ZXt==
j9'AB4,hình 3.1
04^=Y=04';D<,[/(&KbbSchizochytrium>&ZY
s†ZXt=
•Hình 3.1 /k;D<,4# L/! i•;
!=*[7S. AN"•=$/;'Yv]WX9/
;D<,VI;lAH ) I@ 
L/ =K! i;D<,[•E6
l<Fk!,<7I! i=
+Q;•AC[/[FS;Šf=04I,
I,4//,7/WZv 2./C

/,42G7pB7H.</ ;'=
VTF:;&'%Z(e=?Z(eZXD]#SchizochytriumFGH
+SSEIn'< EVVk<AB[/'/
#9/%'7D =b)[AB
k<WXX#w762Wv
X
*4/ WUX < A
N"•=K, 2.[;pS%<AB;
 <AB<7IH[Schizochytrium>&ZY=
+4&A Y~
Khóa luận tốt nghiệp 2014 Khoa Sinh Học
j9'bảng 3.1
K'^=Y=0 <AB<7Iq.%<AB;[/(&KbbSchizochytrium
>&ZY7<)7@h1g>&[1/ +?
] '%Z(e=? Z(eZXDr$s
Schizochytrium >&Y^ ~=U` WU={X
Schizochytrium >&YZ ~={{ Yv=W{
Schizochytrium FGH HVtG VGmu
Schizochytrium >&ZW YY=XU Wv=`^
Schizochytrium >&UY YY=~` Y~=Z`
-/bảng 3.1k/^`k%<AB;AB[/(&Kbb
Schizochytrium >&ZY&j%<AB;.<H[#<,=b
(&Kbb Schizochytrium >&ZY! V'k;.H<AB<H=
VT(h]#:;<=>?@Z+W*W&'(h]#]
1 Schizochytrium FGH
j'VAC[/'FA:))/!?'AC
C!; A;/=12702 i===<.'
'AC;'VAC[/'=
VTH(h]#X
70':AC[/'=g|<,'! 27, 70

ACk?70AC.k=70I'70@
W=U]~7<H';'70.k;'=b)
E'AC[/70AC[/(&KbbSchizochytirum
>&ZYHI'70@Za~=
™AC[/70<;'VAC  , ;.[/(&Kbb
Schizochytirum>&ZYABSCbảng 3.2 hình 3.2
K'^=W=™AC[/70:AC[/ Schizochytrium >&ZY
70
.<AB
s†YX
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w <t &jsw<t
Z ^v=^U v=U`
+4&A WX

×